The anatomy of the nervous system, from the standpoint of development and function . which 208 THE NERVOUS SYSTEM almost at once becomes myelinated and runs through the granular layer into thewhite substance of the cerebellum. According to Clarke and Horsley (1905) andCajal (1911) these axons end in the central cerebellar nuclei. Near their originthey give off collaterals, which run backward through the molecular layer toend in connection with neighboring Purkinje cells—an arrangement designedto bring about the simultaneous discharge of a whole group of such granular layer, situate
The anatomy of the nervous system, from the standpoint of development and function . which 208 THE NERVOUS SYSTEM almost at once becomes myelinated and runs through the granular layer into thewhite substance of the cerebellum. According to Clarke and Horsley (1905) andCajal (1911) these axons end in the central cerebellar nuclei. Near their originthey give off collaterals, which run backward through the molecular layer toend in connection with neighboring Purkinje cells—an arrangement designedto bring about the simultaneous discharge of a whole group of such granular layer, situated immediately subjacent to that which we havejust described, is characterized by the presence of great numbers of small neurons,the granule cells. Each of these contains a relatively large nucleus, surroundedby a small amount of cytoplasm; and from each there are given off from threeto five short dendritic branches with claw-like endings. These are synapticallyrelated with the terminal branches of the moss fibers, soon to be described, and Purkinje cell Basket cell Granule cell. Purkinje cell~ Granule cell Fig. 151.—Diagrammatic representation of the structure of the cerebellar cortex as seenin a section along the axis of the folium (on the right), and in a section at right angles to the axisof the folium (on the left). form with them small glomeruli comparable to those of the olfactory bulb (). Each granule cell gives origin to an unmyelinated axon, which extendstoward the surface of the folium and enters the molecular layer. Here it dividesin the manner of a T into two branches. These run parallel to the long axis ofthe folium through layer after layer of the dendritic expansions of the Purkinjecells, with which they doubtless establish synaptic relations (Fig. 151). Besidesthe granules just described, this layer contains some large cells of Golgis TypeII (Fig. 150, /). Most of these are placed near the line of Purkinje cells andsend their dendrites into the molecu
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