. Asiatic herpetological research. Reptiles -- Asia Periodicals; Amphibians -- Asia Periodicals. Vol. 3, p. 70 Asiatic Herpetological Research April 1990. FIG. 1. Testudo graeca nikolskii is a Mediterranean species, and in the Colchis it is found only in the seaside strip of land with enclaves of Mediterranean vegetation. 2. The Caucasian group includes Hyla arborea schelkownikowi (Fig. 2), Rana macrocnemis, Lacerta caucasica alpina, L. rudis, and Vipera dinniki. Distributions of these species in the Caucasian Isthmus are broader than those of the Colchis group. At the same time, the majority
. Asiatic herpetological research. Reptiles -- Asia Periodicals; Amphibians -- Asia Periodicals. Vol. 3, p. 70 Asiatic Herpetological Research April 1990. FIG. 1. Testudo graeca nikolskii is a Mediterranean species, and in the Colchis it is found only in the seaside strip of land with enclaves of Mediterranean vegetation. 2. The Caucasian group includes Hyla arborea schelkownikowi (Fig. 2), Rana macrocnemis, Lacerta caucasica alpina, L. rudis, and Vipera dinniki. Distributions of these species in the Caucasian Isthmus are broader than those of the Colchis group. At the same time, the majority of them are Colchis autochthons. These species are mesophiles and occur in mesophillous forest and mountain meadow formations. This group seems to be of Colchis origin, retaining close connections with the main center of the formation. Broader ecological tolerance in comparison with typical Colchis species makes it impossible to include them within the Colchis group. 3. The European group consists of Bufo viridis, Rana ridibunda, Emys orbicularis, Anguis fragilis, Natrix natrix, Coronella austriaca, and Coluber jugularis caspius. The composition of this group is not homogeneous. It includes both species typical for the steppe areas {Bufo viridis and Coluber jugularis ) and those that are widely distributed in Europe (all the rest). Only Anguis fragilis and Coronella austriaca are widely distributed in the Colchis. This makes it difficult to definitely consider them late migrants to the Caucasus. Other species either occur in several spots along the Colchis seashore (B. viridis and Natrix natrix ) or populate a narrow strip of land along the sea together with the Mediterranean species (C. jugularis ) or a somewhat wider strip (Emys orbicularis and Rana ridibunda ). Despite the possibility of finding these species in the typical Colchis forest formations, the majority of them are still attributed to the Mediterranean type of Please note that these images are extracte
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