. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. Bonner zoologische Beiträge 55 (2006) 313 gion of the posterior cephalic shield. The Nclc (Nervus clypei capitis) innervates the largest hind part of the pos- terior cephalic shield. We could not detect a lip organ (Fig. 2B), which according to Edlinger (1980) should comprise two small lobes on the cephalic shield above the mouth. A Hancock's organ described by Edlinger (1980) for .4. Tornatilis, here a folded structure separated from the cephalic shield was likewis
. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. Bonner zoologische Beiträge 55 (2006) 313 gion of the posterior cephalic shield. The Nclc (Nervus clypei capitis) innervates the largest hind part of the pos- terior cephalic shield. We could not detect a lip organ (Fig. 2B), which according to Edlinger (1980) should comprise two small lobes on the cephalic shield above the mouth. A Hancock's organ described by Edlinger (1980) for .4. Tornatilis, here a folded structure separated from the cephalic shield was likewise not found in the present study. Tracing studies By conducting the axonal tracing studies we were able to reconstruct cellular innervation patterns for the four cere- bral nerves of A. tonmtilis. Ten replicate tracings were per- formed each for the Nl N2, N3 and Nclc using only the nerves of the right cerebral ganglion. The characteristic patterns of labelled somata for all nerves are shown in Fig- ure 3A-D, including the approximate pathways of the stained axons. The identified clusters were named with ab- breviations signifying the ganglion in which they are lo- cated, the nerve filled and a number indicating the order of their description (for example, Cnlc3: Cerebral Nervus labialis cluster 3). Nerve cells are grouped in clusters on the basis of their close positioning in the ganglia and the tight fasciculation of their axons projecting into the filled nerve. Asymmetries for tracings of the left nerves could not be detected. For the Nl (n = 10) we identified six cerebral clusters (Cnocl-6) and one pedal cluster (PdnocI) in each sam- ple (Fig. 3A). The variation between the samples was re- stricted to very few somata in some clusters. The cerebral clusters were distributed over the whole cerebral ganglion. The pedal cluster PdnocI was located on the anterior mar- gin of the pedal ganglion above the pedal commissure. The innervation pattern of the N2 (n=10) consists o
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