. Bulletin. Science; Natural history; Natural history. FLOOD EFFECTS ON INTERTIDAL BIOTAS 101 ,1^^. y^r^<^^;<:<;^'" ^^^^^^...^^^ <:;x o<X ^*^' aN ,1^' pHf^^' CO^' 5^f^^ Fig. 4. Macroinvertebrate cover (features are the same as those given in Fig. 3). disturbance gradients relative to the maintenance of species diversity. However, the phenomenon documented here is relatively species selective, although pre- sumably not an isolated occurrence (cf. Sousa et al. 1981). It would have been optimal if we had been able to compare the transect area affected by the flood with an iden
. Bulletin. Science; Natural history; Natural history. FLOOD EFFECTS ON INTERTIDAL BIOTAS 101 ,1^^. y^r^<^^;<:<;^'" ^^^^^^...^^^ <:;x o<X ^*^' aN ,1^' pHf^^' CO^' 5^f^^ Fig. 4. Macroinvertebrate cover (features are the same as those given in Fig. 3). disturbance gradients relative to the maintenance of species diversity. However, the phenomenon documented here is relatively species selective, although pre- sumably not an isolated occurrence (cf. Sousa et al. 1981). It would have been optimal if we had been able to compare the transect area affected by the flood with an identical control area unaffected by the flood (both before and after). However, since this was not possible, we used (1) an adjacent unflooded control area (visually similar) established after the flood, (2) a com- parable but not identical control area 40 km to the north ( Whites Point, Murray and Littler 1984), and (3) the previous two years of detailed seasonal data at Corona del Mar in which no flooding occurred. Our method of nondestructive sampling, by utilizing the identical sampling locations that were precisely relocated and reassessed, provided a powerful tool for the quantification of natural changes in the intertidal standing stocks, because variance due to sampling design was virtually eliminated. Throughout the two years of quarterly sampling at this site prior to the flood, seasonal patterns tended to be both minor and predictable (Littler 1977, 1978). Historically, plant cover at Corona del Mar increased annually during spring to late summer followed by slight declines in late fall and early winter associated with aerial exposures during daytime low tides and some rock tumbling due to storm waves. Animal cover typically showed little seasonal change, except for vertical migratory movements of mobile limpet species (Seapy and Hoppe 1973) and a sporadic winter to spring recruitment of barnacles (Littler 1980a), that appeared to be minimally affected by the flood
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