. Bulletin of the British Museum (Natural History), Geology. 162 P. E. S. WHALLEY R,b *i° Jl. Fig. 60 Archaeolepis mane gen. et sp. nov. (Lepidoptera). Composite diagram of wing venation. A = anal veins, Cu = cubital vein, M = median veins, R = radial vein, Rs = branches of radial sector, Sc = subcostal vein. the longitudinal ridges are apparent near base of some scales. Apex of scales smoothly rounded. Two, possibly more, different types of scales. Marginal scales thinner, some thinner scales mixed in with thicker ones on wing membrane. The wing margin has longer scales along the hind margin
. Bulletin of the British Museum (Natural History), Geology. 162 P. E. S. WHALLEY R,b *i° Jl. Fig. 60 Archaeolepis mane gen. et sp. nov. (Lepidoptera). Composite diagram of wing venation. A = anal veins, Cu = cubital vein, M = median veins, R = radial vein, Rs = branches of radial sector, Sc = subcostal vein. the longitudinal ridges are apparent near base of some scales. Apex of scales smoothly rounded. Two, possibly more, different types of scales. Marginal scales thinner, some thinner scales mixed in with thicker ones on wing membrane. The wing margin has longer scales along the hind margin and probably had a substantial fringe of scales. Fewer ridges on some scales, 12-13 as against 18-22 on others. The ridges on the scales as seen in the stereoscan micro- graphs (Fig. 58d) are considered to represent the infilling between the original ridges (striae) and their compression, with the dark lines representing the ridges from the original scale. The scales shown in the micrographs are the narrow ones with fewer striae. Holotype. (Figs 58-59); Birchi nodules, Lower Lias, Black Ven, Charmouth, Dorset; part and counterpart. Jackson colln. Dimensions: Length 5-3-5-6mm, width 2mm. Discussion. Although Lepidoptera or Trichoptera are the most obvious orders for the fossil wing, others cannot be eliminated without further consideration. Indeed some of the Recent scaly-winged Lepidopsocidae (Psocoptera) look superficially very similar to the Dorset fossil. Apart from these, consideration was also given to the possibility that the wing represented an extinct, scale-winged mecopteran (sens. lat.). Several families in the broad assemblage Meco- pteroidea (Willmann 1978) have been considered as giving rise to the modern Diptera, Tricho- ptera and Lepidoptera (Tillyard 1933). Many Jurassic Orthophlebiidae have an unbranched cubital vein, contrasting with Archaeolepis, and are really early Mecoptera. The Pseudo- polycentropodidae, which have a highly specialized and sligh
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