. Current herpetology. Reptiles; Herpetology. 80 Current Herpetol 21(2) 2002 (r=, P= at Bonbori; r=, P= at Tochigi). All females oiR. sakuraii but one deposited eggs of more than 3 mm in diameter, and most plots of R. sakuraii were located above those of R. t. tagoi (Fig. 4). These results indicate that the egg size of R. sakuraii was larger than that of R. t. tagoi, even if adjusted for body size, and that the mean egg size of the R. sakuraii population was significantly larger at Tochigi than at Bon- bori (t=, in both), but differed significantly in elevation b


. Current herpetology. Reptiles; Herpetology. 80 Current Herpetol 21(2) 2002 (r=, P= at Bonbori; r=, P= at Tochigi). All females oiR. sakuraii but one deposited eggs of more than 3 mm in diameter, and most plots of R. sakuraii were located above those of R. t. tagoi (Fig. 4). These results indicate that the egg size of R. sakuraii was larger than that of R. t. tagoi, even if adjusted for body size, and that the mean egg size of the R. sakuraii population was significantly larger at Tochigi than at Bon- bori (t=, in both), but differed significantly in elevation between populations oi R. sakuraii and R. t. tagoi (P< in all combinations). The adjusted means of the clutch volumes were 1727, 1694, 2041, and 2080 mm^ for the Bonbori, Tochigi, Ichi- nosawa, and Kamiange populations, respec- tively (Fig. 5). Clutch volume of R. sakuraii was, therefore, approximately 17% smaller 4000. 40 50 60 SNOUT-VENT LENGTH (mm) Fig. 5. Relationship between female body size and clutch volume. The log-log regression did not differ significantly between two populations in each species, and a single regression equation was thus calculated from combined conspecific data sets. Solid and dashed lines indicate regressions of R. sakuraii (logY=, r=, n=55) and R. t. tagoi (logY=, r=, n=52), respectively. than that of R. t. tagoi when compared between similar-sized females. Age and clutch parameters A total of 26 females of R. t. tagoi (18 and eight from Ichinosawa and Kamiange, respec- tively) was aged by skeletochronology using phalanges. Their ages ranged from two to four years (see also Kusano et al., 1995a). Because clutch parameters were closely related to body size as mentioned above (Figs. 3-5), it was necessary to examine the relationship between age and clutch parameters after remov- ing the effect of female SYL. Relative egg and clutch sizes were, therefore, calculated as residuals from the regressions of the


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