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. The Biological bulletin. Biology; Zoology; Marine biology. COLONY Figure 4. Means and standard errors of rates times 10' of lumen expansion and contraction (/im of lumen diameter per total ^m of stolon width per second) for 10 colonies of each of P. cornea (open bars) and H. symbiolongicarpus (filled bars) at three stages of colony development (a = stage 1, b = stage 2, c = stage 3). Colonies of the two species are paired arbitrarily. (paired comparison Mest of mean rates for each colony, / = , P < ) and between stage 1 and 3 (t = —, P < ). This decrease in rate is br
. The Biological bulletin. Biology; Zoology; Marine biology. COLONY Figure 4. Means and standard errors of rates times 10' of lumen expansion and contraction (/im of lumen diameter per total ^m of stolon width per second) for 10 colonies of each of P. cornea (open bars) and H. symbiolongicarpus (filled bars) at three stages of colony development (a = stage 1, b = stage 2, c = stage 3). Colonies of the two species are paired arbitrarily. (paired comparison Mest of mean rates for each colony, / = , P < ) and between stage 1 and 3 (t = —, P < ). This decrease in rate is brought about by a decrease in the mean relative amplitude between stage 1 and 2 (t = , P < ) and stage 1 and 3 (t = , P ) and between stage 1 and 3 (t = , P < ). Interestingly, the variability of the sto- lonal contraction rate (including both within- and be- -tween-stolon variance) significantly decreases between stage 1 and 2 (paired comparison r-test of the standard deviations of rates for each colony, t = — \,P ). Thus the rate of stolonal expansion and contraction in H. sym- biolongicarpus shows the least variability at the time of stolonal mat formation (Fig. 5; see discussion in Black- stone and Buss, 1993), although the actual mean rate is lowest at stage 3. The change in the variability of rate is brought about largely by changes in the variability of rel- ative amplitude, the standard deviation of which decreases significantly between stage 1 and 2 (t = —, P ). Cycle period shows no change in variability (between stage 1 and 2, t = , P > ; between stage 1 and 3, t = , P> ). In P. carnea, mean rate significantly increases between stage 1 and 2 (paired comparison Mest of mean rates for each colony, t = , P < ) and between stage 1 and 3 (? = , P <^ ). This increase in rate is brought about by an increase in the mean relative amplitude be- tween stage 1 and 2{t= , P <
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