. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. 172 A. Tye and sympatry (in Transcaucasia and the Mangyshlak Peninsula: Loskot 1986; Panow 1986) but Haffer (1977) considered the degree of introgression insufficient to warrant merging the two into a single species. The present superspecies appears to stand between species which are mainly black- and-white and those which are mainly sandy-brown. It is perhaps most closely- related to the O. picata species-group: male pleschanka in worn dress closely resemble O. lug


. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. 172 A. Tye and sympatry (in Transcaucasia and the Mangyshlak Peninsula: Loskot 1986; Panow 1986) but Haffer (1977) considered the degree of introgression insufficient to warrant merging the two into a single species. The present superspecies appears to stand between species which are mainly black- and-white and those which are mainly sandy-brown. It is perhaps most closely- related to the O. picata species-group: male pleschanka in worn dress closely resemble O. lugens, a member of the picata group. Fig. 4: Breeding ranges of O. picata species-group. Numbers indicate subspecies of O. lugens as follows: 1 halophila, 2 lugens, 3 pérsica, 4 lugentoides, 5 boscaweni, 6 lugubris, 1 vauriei, 8 schalowi. I include here the Eastern Pied Wheatear O. picata, the Mourning Wheatear O. lugens, and Finsch's Wheatear O. finschii. O. picata and O. lugens have previously been associated as a superspecies (Mayr & Stresemann 1950; Hall & Moreau 1970), or even as members of a single species (Stresemann 1925), while O. finschii has been regarded as a subspecies of O. lugens (e. g. by Dementiev et al. 1954). In fact, the breeding ranges of O. lugens and O. finschii overlap in Jordan and the Zagros Moun- tains of western Iran (Fig. 4), so these two are good species, although evidently closely-related (see Cramp 1988). Further, the range of O. picata overlaps those of lugens in eastern Iran and finschii in northern Iran and Afghanistan (Fig. 4). O. picata is one of the most variable species of wheatear (see Mayr & Stresemann 1950, Panow 1980), while O. lugens is certainly the most polytypic and also shows polymorphism in some of its races. In contrast O. finschii is not polymorphic (although it has sexual dimorphism) and geographical variation in it is slight (see Ticehurst 1927). O. picata has three major plumage types, capistrata, opistholeuca an


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