. Bulletin of the British Museum (Natural History), Geology. 10. Fig. 5 Laveineopteris rarinervis (Bunbury) Cleal et al. Copy of Carpentier (1930: fig. 1), showing orbicular cyclopterids attached near the dichotomy of the primary rachis. Here reproduced at x 0-7 life size. with the Trigonocarpales (Oestry-Stidd 1979). Little is known of the fructifications, other than that large ovules were attached directly to the frond, probably at the end of ultimate pinnae (Kidston 1904). Form-genus MACRONEUROPTERIS Cleal, Shute & Zodrow (1990: 488) Type. M. macrophylla (Brongniart) Cleal, Shute &
. Bulletin of the British Museum (Natural History), Geology. 10. Fig. 5 Laveineopteris rarinervis (Bunbury) Cleal et al. Copy of Carpentier (1930: fig. 1), showing orbicular cyclopterids attached near the dichotomy of the primary rachis. Here reproduced at x 0-7 life size. with the Trigonocarpales (Oestry-Stidd 1979). Little is known of the fructifications, other than that large ovules were attached directly to the frond, probably at the end of ultimate pinnae (Kidston 1904). Form-genus MACRONEUROPTERIS Cleal, Shute & Zodrow (1990: 488) Type. M. macrophylla (Brongniart) Cleal, Shute & Zodrow Comments. Most neuropteroid fronds have a dichotomy of the primary rachis producing tripinnate or occasionally quad- ripinnate branches. In some species, however, the dichotomy of the primary rachis produces less-divided, essentially bipin- nate, primary rachis branches (Figs 6-7). These species also have a number of distinctive epidermal characteristics, such as brachyparacytic or cyclocytic stomata (Fig. 17E,F). It was for this distinctive group of species that Cleal et al. (1991) proposed the form-genus Macroneuropteris. The fronds of Macroneuropteris are very similar to Neurop- teris sensu stricto, except that they are less divided. Of particular significance is the presence in at least one macro- neuropterid species (M. scheuchzeri) of so-called 'Odontopt- CLEAL AND SHUTE eris lindleyana'- type pinnules ( Crookall 1959: pi. 57, fig. 1), which can be compared with laciniate pinnules in the lower part of true neuropterid fronds ( Stockmans 1933: pi. 11 fig. 1; pi. 12 fig. 2; Zodrow & Cleal 1988: pi. 4 fig. 3). There is no evidence of the orbicular cyclopterid pinnules of Laveineopteris or Margaritopteris. Nothing is known of the fructifications. Beeler (1983) claimed that the rachis anatomy is of a type typical of the Trigonocarpales. However, this was based purely on evi- dence of association; she could find no such rachides with macroneuropteri
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