. Bulletin of the Museum of Comparative Zoology at Harvard College. Zoology. 146 Bulletin Museum of Comparative Zoology, Vol. 145, No. 2 i, spermotheco. HIRTUS CLUSTER JLOEDINGI "cluster" HAPARDI OCYTUS FISUS SCHWARZI TEXANUS CALIFORNICUS â | |G1AQUINT01 EVADICUS CONSOBRINUS BREVIOR ULKEI T-Q CAVERN I COLA '-QcAVERNICOLA ADITUS GYPSUM troglomexicahus Qelabra JAME5I ALTUS NEWTON I â QSPELAEUS LEO 3 C o T ertiary Ouater nary CO TO a> Q 3 â IQ _.(t> _. =j o â ^ i^ =" 3 "-* 3 ZT 2-° 5 cave occupation 213 Figure 213. Hypothetical phylogenetic diagram of specific diversifi


. Bulletin of the Museum of Comparative Zoology at Harvard College. Zoology. 146 Bulletin Museum of Comparative Zoology, Vol. 145, No. 2 i, spermotheco. HIRTUS CLUSTER JLOEDINGI "cluster" HAPARDI OCYTUS FISUS SCHWARZI TEXANUS CALIFORNICUS â | |G1AQUINT01 EVADICUS CONSOBRINUS BREVIOR ULKEI T-Q CAVERN I COLA '-QcAVERNICOLA ADITUS GYPSUM troglomexicahus Qelabra JAME5I ALTUS NEWTON I â QSPELAEUS LEO 3 C o T ertiary Ouater nary CO TO a> Q 3 â IQ _.(t> _. =j o â ^ i^ =" 3 "-* 3 ZT 2-° 5 cave occupation 213 Figure 213. Hypothetical phylogenetic diagram of specific diversification in Ptomajphagus [Adelops]. Degree of specific difference not inferred. Rates of divergence in Tertiary not inferred. groups in the subgenus Adelops upon the female spermatheca, then- characters and complexities. This is the only character I found that provided me with data I could use in interpreting a phylogeny for major events within the subgenus. In the discus- sion to follow only North and Central American events will be considered. I want to caution here against a too ready acceptance of the chronologic impli- cations of the following discussion. Where I suggest, for instance, that two related events occurred in the Illinoian and Wisconsin respectively, more definitely "earlier" and "later" are being indicated. This is based upon the assumption that greater differences take a longer time to develop than lesser ones. I offer definite times only as appealing possibilities that seem to be logically consistent within an overall framework of combinations of fact, theory, and circumstantial evidence. Figure 213 presents a summary of the following discussion. I believe the hirtus group to be the most primitive, and I base this belief solely upon the observation that within this group the female spermatheca is the most simple, and is closest in fonn to the other genera in the Ptomaphagina (Echinocoleus and Adelopsis). The hirtus group is hence the


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