. Bonn zoological bulletin. Zoology. 4 Johan van Rooijen et Fig. 1. A: View into the primary forest of Pangkor Island; B: Boiga drapiezii (Boie, 1827); C: Cnemaspis shahruli Grismer et al., 2010; D: Diyophiops rubescens (Gray, 1835). island is Pulau Pangkor, approximately 18 km2 and situ- ated km from the west coast of Peninsular Malaysia. The island's interior remains heavily forested and main- tains a river system, Sungai Pinang, which supplies a sig- nificant source of permanent fresh water in the form of multiple streams. Chan et al. (2010) provided the first re- port on the (non-


. Bonn zoological bulletin. Zoology. 4 Johan van Rooijen et Fig. 1. A: View into the primary forest of Pangkor Island; B: Boiga drapiezii (Boie, 1827); C: Cnemaspis shahruli Grismer et al., 2010; D: Diyophiops rubescens (Gray, 1835). island is Pulau Pangkor, approximately 18 km2 and situ- ated km from the west coast of Peninsular Malaysia. The island's interior remains heavily forested and main- tains a river system, Sungai Pinang, which supplies a sig- nificant source of permanent fresh water in the form of multiple streams. Chan et al. (2010) provided the first re- port on the (non-marine) herpetofauna of this island and documented 43 reptiles and 13 amphibians. A few illus- trations are provided in figures 1 and 2. The study described in this paper had a dual objective. First, data collected by Chan et al. (2010) were used to evaluate the performance of several estimators. Second, the most appropriate estimator was used to estimate the total herpetofaunal species richness harboured by Pangkor Island, West Malaysia. MATERIAL AND METHODS The data underlying this study were based on the surveys carried out by Chan et al. (2010). These were conducted in the periods March 15-17, May 4-8, June 13 to July 8, 2009, and February 22 to March 8, 2010. During the lat- ter two periods, only reptiles were surveyed. Marine rep- tiles were ignored altogether. The predominantly applied survey method corresponds to visual encounter survey (VES), a simple method which has been shown to be ef- fective for surveying rainforest herpetofauna (Doan, 2003). VES was earned out both during day and night ( Coddington et al., 1996; Doan, 2003) in a way similar to that applied by Minh (2007). Existing trails as well as sev- eral trails made by the Department of Forestry of Perak and Peninsular Malaysia were used. These trails traversed dipterocarp forest, mangrove forest and cultivated areas and provided ample access to forest streams. The second collection method consisted of se


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