. Bonn zoological bulletin. Zoology. Boophis quasiboehmei sp. n., a new cryptic treefrog species from south-eastern Madagascar 247 # Boophis boehmei O Boophis quasiboehmei. PCA Factor 2 Fig. 3. Scatterplot of individual males of Boophis boehmei (filled circles) and B. quasiboehmei (open circles) along the sec- ond and third factor of a Principal Component Analysis (Vari- max normalized rotation). The PCA was based on measurements in Table 1. The specimen from Midongy was excluded from analysis because the species identity of this population
. Bonn zoological bulletin. Zoology. Boophis quasiboehmei sp. n., a new cryptic treefrog species from south-eastern Madagascar 247 # Boophis boehmei O Boophis quasiboehmei. PCA Factor 2 Fig. 3. Scatterplot of individual males of Boophis boehmei (filled circles) and B. quasiboehmei (open circles) along the sec- ond and third factor of a Principal Component Analysis (Vari- max normalized rotation). The PCA was based on measurements in Table 1. The specimen from Midongy was excluded from analysis because the species identity of this population is not ful- ly clarified. al steps. Although the nuclear data set refers to only a lim- ited number of specimens, the fact that there is no haplo- type sharing between the two forms suggests that they rep- resent independent evolutionary lineages. Nevertheless, these pronounced genetic divergences were contrasted by no or low divergences in adult morpholo- gy and bioacoustics. The calls of the two forms were sim- ilar, with no detectable differences (see call descriptions below). In both forms, notes may be combined to short regular series, and intervals between notes are otherwise highly variable and mostly irregular. The temporal and spectral parameters in calls of both forms are somewhat variable among populations and individuals, but broadly overlap at inter- and intra-populational level. Even the pulse rate within notes, a character shown to be evolution- ary highly dynamic among closely related species ( Pa- dial et al. 2008), is identical in both forms (see analysis below). Inter-note intervals outside of regular note series furthermore seem to depend on calling motivation of the individual male. A close examination of adult morphology yielded no dis- crete characters that would allow a diagnosis between the two forms. One subtle difference was detected in adult life A C E F ^^^^^^^^^^^^^ Fig. 4. Comparative photographs of oral discs of preserved tadpoles of Boophis
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