. The Ecology of arboreal folivores : a symposium held at the Conservation and Research Center, National Zoological Park, Smithsonian Institution, May 29-31, 1975 . Figure 1. Approximate locations of 2 study areas on Barro Colorado Island. Area A is Lutz Ravine near laboratory clearing. Area B is Old Forest. the eastern side of the island, was selected because it had been the site of all previous detailed investigations of howler behavior on Barro Colorado Island (Car- penter, 1934; Collias and Southwick, 1952; Altmann, 1959; Bernstein, 1964; Chivers, 1969; Hladik and Hladik, 1969; Richard, 19
. The Ecology of arboreal folivores : a symposium held at the Conservation and Research Center, National Zoological Park, Smithsonian Institution, May 29-31, 1975 . Figure 1. Approximate locations of 2 study areas on Barro Colorado Island. Area A is Lutz Ravine near laboratory clearing. Area B is Old Forest. the eastern side of the island, was selected because it had been the site of all previous detailed investigations of howler behavior on Barro Colorado Island (Car- penter, 1934; Collias and Southwick, 1952; Altmann, 1959; Bernstein, 1964; Chivers, 1969; Hladik and Hladik, 1969; Richard, 1970; Mittermeier, 1973; Smith, 1977). I felt that a long-term study in this area would help to confirm and tie together many of the scattered pieces of information on howlers in the Lutz Ravine. The second study site, which I call the Old Forest, was located some 45 minutes by foot from the laboratory clearing in an area which is roughly bor- dered by the junctions of the Wheeler-Drayton and Zetek-Armour trails. The forest in Lutz Ravine is largely composed of trees less than 100 years old (Bennett, 1963; Foster, personal communication). Much of this area is known to have been cleared some 60 to 80 years ago before Barro Colorado was established as a nature preserve. This apparently allowed the growth of a large num- ber of nonstrangler (Pharmacosycea) fig (Ficus) trees which typically begin growth only in relatively open areas. There are approximately 127 large nonstrangler fig trees of this type in the 25 ha making up the Lutz Ravine. Quantitative data show that this is about 9 times as many fig trees per hectare as are generally found on the island (Knight, 1975). The second study site, the Old Forest, has in large part been undisturbed for at least the past 130 to 150 years and is believed to be structurally mature (Foster, personal communication). There are patches of younger secondary forest in this area which the troop regularly included as part of its home range. Most no
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Keywords: ., bookcentury1900, bookcolle, booksubjectleaves, booksubjectmammals
