. Annals of the South African Museum = Annale van die Suid-Afrikaanse Museum. Natural history. Fig. 14. Undamaged (above) and acid-etched (below) penguin tibiotarsi, femora and carpometacarpi from the QSM (East Stream sample). the case of the 'E' Quarry penguins. In addition, penguins are preyed upon by terrestrial animals, and 'E' Quarry is unique amongst recorded Tertiary pen- guin localities in that the penguin bones come from non-marine deposits (Simpson 1976) occurring in association with a wide variety of terrestrial vertebrates, including predaceous forms. Consequently, it could also be


. Annals of the South African Museum = Annale van die Suid-Afrikaanse Museum. Natural history. Fig. 14. Undamaged (above) and acid-etched (below) penguin tibiotarsi, femora and carpometacarpi from the QSM (East Stream sample). the case of the 'E' Quarry penguins. In addition, penguins are preyed upon by terrestrial animals, and 'E' Quarry is unique amongst recorded Tertiary pen- guin localities in that the penguin bones come from non-marine deposits (Simpson 1976) occurring in association with a wide variety of terrestrial vertebrates, including predaceous forms. Consequently, it could also be signi- ficant that 'E' Quarry is apparently the only known locality from which penguin bones etched by stomach acids have been recorded. In fact, the 'E' Quarry penguins are likely to have been preyed upon by several aquatic and terrestrial predators, although only one is likely to have caused the acid-etched bones. This was probably an animal that bolted its food, and subsequently regurgitated indigestible residues. This behaviour applies in the case of sharks (P. A. Hulley, pers. comm.), and of the predators recorded in association with the 'E1 Quarry penguins, sharks are here favoured for the role. Further clues concerning predation on the 'E' Quarry penguins may emerge from more detailed examinations of the available material and compari- sons with stomach contents of modern penguin predators. Even the most superficial examination of the 'E' Quarry penguin assemblages reveals features that may be of taphonomic significance. For example, etching is more pro- nounced on leg bones than those of the flipper (Table 5). In addition, certain. Please note that these images are extracted from scanned page images that may have been digitally enhanced for readability - coloration and appearance of these illustrations may not perfectly resemble the original South African Museum. Cape Town : The Museum


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Keywords: ., bookcentury1800, bookdecade1890, booksubjectnaturalhistory, booky