. Fungi, ascomycetes, ustilaginales, uredinales. Fungi. 58 PLECTOMYCETES [CH. Only two species of Eremascus are known. E. albtis was discovered by Eidam in 1881, in a bottle of malt extract. The contents had gone bad and their surface was covered with a growth of various fungi, amongst which was the new genus. It pro- duced a fine, snowy white, septate mycelium from which pairs of fer- tile hyphae grew out, curled round one another and fused at their tips (fig. 18). The fused portion was cut off from the fertile hypha below, and eventually produced eight spores. Unfortunately Eidam's species w
. Fungi, ascomycetes, ustilaginales, uredinales. Fungi. 58 PLECTOMYCETES [CH. Only two species of Eremascus are known. E. albtis was discovered by Eidam in 1881, in a bottle of malt extract. The contents had gone bad and their surface was covered with a growth of various fungi, amongst which was the new genus. It pro- duced a fine, snowy white, septate mycelium from which pairs of fer- tile hyphae grew out, curled round one another and fused at their tips (fig. 18). The fused portion was cut off from the fertile hypha below, and eventually produced eight spores. Unfortunately Eidam's species was lost and has never reappeared. In 1907, however, Stoppel, on opening some pots of apple and gooseberry jelly, discovered a very similar form which she named Ere- mascus fertilis. This species, like E. albus, possesses a branching, septate mycelium. The cells at first contain several nuclei; these, according to Stoppel, are arranged in pairs, but Guilliermond, in a subsequent investigation, found that such an arrangement, even when present in the young mycelium, did not persist. It is no doubt dependent upon rapidity of grcrwth. From this mycelium pairs of uninucleate branches grow up, usually from the same, sometimes from different hyphae, and fuse at their apices (fig. 19). Their nuclei also fuse and after three karyokinetic divisions eight spores are formed. Sometimes, especially in old cultures, the fertile hyphae may produce asci without fusion. These are usually small and generally contain four spores or even a lesser number. As a rule three nuclear divisions take place in the parthenogenetic asci, and eight nuclei are formed, though they do not all function. According to Guilliermond it would seem that the number of spores is conditioned not by any cytological peculiarity, but rather by the supply of nutritive material. The species of the genus Endomyces possess a branched, septate myce- lium. It may break up into oidia, which sometimes become surrounded by thick walls
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