. Ecuadorian frogs of the genus Colostethus (Anura:Dendrobatidae) . Fig. 1. Colostethus chocoensis. A. Right hand (MNHG (GO 47]). B. Left foot (MNHG 19107). Notice webbing between fingers on hand. Foot illustrates a webbing formula of II — l'/illl — l'/2llll—2IV2—IV. Lines equal 1 mm. ter, whereas others (, C. bocagei, C. chocoensis, , and C. nexipus) well-developed fringes on hands, but the fringes on the feet are continuous with the interdigital webbing. Most spe- cies lack fringes; in some, such as C. awa, C. kingsburyi, C. mystax, and C. shuar, fringes are poorly deve
. Ecuadorian frogs of the genus Colostethus (Anura:Dendrobatidae) . Fig. 1. Colostethus chocoensis. A. Right hand (MNHG (GO 47]). B. Left foot (MNHG 19107). Notice webbing between fingers on hand. Foot illustrates a webbing formula of II — l'/illl — l'/2llll—2IV2—IV. Lines equal 1 mm. ter, whereas others (, C. bocagei, C. chocoensis, , and C. nexipus) well-developed fringes on hands, but the fringes on the feet are continuous with the interdigital webbing. Most spe- cies lack fringes; in some, such as C. awa, C. kingsburyi, C. mystax, and C. shuar, fringes are poorly developed. In some cases, the intraspecihc variation observed may be an artifact of preserva- tion that tightens the skin to the digit. When it was difficult to distinguish between a real fringe and a preservation artifact, I describe the dermal modifi- cation as a "; Third finger in males.—The distal segments of the third finger in males is distinctly swollen in several Colostethus and Epipedobates. Myers et. al (1991) indicated that this character is expressed more strongly in some species, and that in some species at least, it is not expressed in all adult males. Of the Ecuadorian Colostethus, only C. machalillci have this character. Tarsal folds.—There are two kinds of folds. A dermal fold may occur along the outer ventral edge of the tarsus. About half of the Ecuadorian Colostethus lacks this fold; however, it is well
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