. Handbook of flower pollination : based upon Hermann Mu?ller's work 'The fertilisation of flowers by insects' . Fertilization of plants. 48 ANGJOSPERMAE—DICOTYLEDONES below, with only a short lip on its free margin. In A. palmatum Wall, the spur makes its first appearance as a slight projection ; in A. Napellus L. the lip is elongated and expanded; in A. Anthora Z. and A. columbinum NuU. the spur is more prominent, and in A. paniculatum Lam. it is well defined. The spur of the Japanese species A. Fischeri Reichmb. is prolonged and curved inwards like a flamingo's bill; in A. septentrionale Ko


. Handbook of flower pollination : based upon Hermann Mu?ller's work 'The fertilisation of flowers by insects' . Fertilization of plants. 48 ANGJOSPERMAE—DICOTYLEDONES below, with only a short lip on its free margin. In A. palmatum Wall, the spur makes its first appearance as a slight projection ; in A. Napellus L. the lip is elongated and expanded; in A. Anthora Z. and A. columbinum NuU. the spur is more prominent, and in A. paniculatum Lam. it is well defined. The spur of the Japanese species A. Fischeri Reichmb. is prolonged and curved inwards like a flamingo's bill; in A. septentrionale Koelle it is drawn out in the shape of a proboscis to a length of 6 mm. ; and lastly, in A. Lycoctonum Z., it is coiled into a spiral of i|- turns. The blossoms of Aconitum are—as Kronfeld expresses it—' humble-bee flowers par excellence' The body of a humble-bee exactly fills the interior of the flower, and if a plaster cast of this be made it will be found to correspond in a remarkable way to the shape of a medium-sized female humble-bee. As a matter of fact, Aconitum is dependent on Bombus, and must become extinct in places where humble-bee visits. Fig. 17. Slaves in Specialisation of the Nectaries of Aconitum (after M. Kronfeld). Nectary of: — I. Aconitum hcterophyllum Wall, (with no spur). 2. A. palmatum Wall, (with a slight projection), 3. A. Napellus L. (with a somewhat larger projection, and a longer lip). 4. A. Anthora L. and A. columbinum L. (with projection still more developed). 5, A. paniculatum Lam. (well-de6ned spur). 6. A. volubile Pall, and E. villosura Rffl. (with buckle-shaped posterior projection, 7, A, Fischeri Reichenb. (spur curved inwards). 8. A. septentrionale Koelle (spur rolled inwards), Q. A. Lycoctonum L. (spur rolled inwards to a greater extent). fail, or where these insects merely rob the flowers by making lateral perforations. The dependence of monkshoods on humble-bees is best made clear by comparing their respective areas of distribution.


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