. Vegetation of Oregon and Washington. Botany Oregon Ecology; Botany Washington (State) Ecology. Figure 113. — Pinus contorta is a common serai spe- cies in the Abies grandis zone of east- ern Oregon; Abies grandis and Picea en- gelmannii succeeding Pinus contorta in the Blue Mountains of Oregon (range pole is marked in 1-foot segments). rossii, Sitanion hystrix, Gayophyturn dif- fusum, Cryptantha affinis, Fragaria cuneifolia, Epilobium angustifolium, Chimaphila umbel- lata, and Apocynum androsaemifolium make up most of the sparse herbaceous layer. Successional Patterns Data on early successio
. Vegetation of Oregon and Washington. Botany Oregon Ecology; Botany Washington (State) Ecology. Figure 113. — Pinus contorta is a common serai spe- cies in the Abies grandis zone of east- ern Oregon; Abies grandis and Picea en- gelmannii succeeding Pinus contorta in the Blue Mountains of Oregon (range pole is marked in 1-foot segments). rossii, Sitanion hystrix, Gayophyturn dif- fusum, Cryptantha affinis, Fragaria cuneifolia, Epilobium angustifolium, Chimaphila umbel- lata, and Apocynum androsaemifolium make up most of the sparse herbaceous layer. Successional Patterns Data on early successional stages following logging or burning on Abies grandis habitats are scarce. Cirsium vulgare was a major species on a clearcut and burned area in the Wallowa Mountains (Pettit 1968); Ceanothus sangui- neus, Physocarpus malvaceus, Spiraea betuli- folia, var. lucida, Astragalus sp., Fragaria spp., Carex rossii, Epilobium paniculatum, Bromus tectorum, and Rumex acetosella were also noted. Later in preforest succession, shrub communities are encountered in the Abies grandis Zone, at least in eastern Washington and northern Idaho (Daubenmire 1952; Mueg- gler 1965). Typical shrub invaders (Dauben- mire and Daubenmire 1968) are Salix scoul- eriana, Spiraea betulifolia var. lucida, Ceano- thus velutinus, C. sanguineus, Amelanchier al- nifolia, Sambucus cerulea, and S. racemosa var. melanocarpa. Mueggler (1965) related the composition of these shrub communites to type of originating disturbance and age. Throughout much of the Abies grandis and Abies concolor Zones in the Cascade Range, Ceanothus velutinus is again a major brush- field dominant, often behaving ecologically as described for southwestern Oregon. Ceano- thus velutinus and species such as Castanopsis chrysophylla and Arctostaphylos patula may form a sclerophyll scrub following wildfires in the central and southern Oregon Abies grandis-A. concolor Zones. As mentioned, Pinus contorta, P. ponder- osa, and Pseudotsuga menzies
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