. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. Fig. 8: Caudal part of right branch of mandíbula of Mirolia? mascalidris spec. nov. in frontal (A), lateral (B) and medial (C) view. Scale 1 mm. 1) long processus retroarticularis 2) big insertion area for m. depressor mandibulae 3) slender processus interior 4) foramen pneumaticum 5) OS complementare joins os suprangulare. thus separating the fontanella from the channel for the n. mandibulare (a) 6) open fontanella 13. RESULTS AND CONCLUSIONS Comparative studies ha
. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. Fig. 8: Caudal part of right branch of mandíbula of Mirolia? mascalidris spec. nov. in frontal (A), lateral (B) and medial (C) view. Scale 1 mm. 1) long processus retroarticularis 2) big insertion area for m. depressor mandibulae 3) slender processus interior 4) foramen pneumaticum 5) OS complementare joins os suprangulare. thus separating the fontanella from the channel for the n. mandibulare (a) 6) open fontanella 13. RESULTS AND CONCLUSIONS Comparative studies had to be carried out on skeleton material of recent Charadriiformes for the determination of the systematic position of the fossils here described from the Miocene of the Nördlinger Ries. The oste- ological data support a separation of the Scolopacidae from the Charadriidae at the family level. This view has already been expressed elsewhere, in the phyloge- netic tree by Strauch (1978: flg. 36). Within the Scolopacidae, osteological differences be- tween Calidridinae and Tringinae are in the first place found on the cranium and the bill. In addition, some complex postcranial structures such as the processus su- pracondylaris dorsalis of the humerus can help to distin- guish the two groups. The bulk of the fossil remains of small Scolopacidae de- scribed here from the middle Miocene of the Nördlinger Ries belongs to several species of Mirolia, a new fossil genus within the Calidridinae. Thus the decision of Brodkorb (1967), who assigned a number of fossils from the Miocene to this subfamily is justified. It appeared that generic determination of Calidridinae of Miocene age can only be based on rather complete fos- sil material: the diagnosis of Mirolia and the more de- tailed discussion of its systematic position rely in first place on cranial features. The nearest relatives of the fossil genus Mirolia might be the small Scolopacidae of La , espe- cially 'E
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