. Bulletin of the British Museum (Natural History) Geology Supplement. FOSSIL AND RECENT ors 15 I so. pto asp. Mot myp Fig. 4. Elops hawaiensis Regan. Neurocranium in orbital view. hyomandibular facet. The orbital face of the autosphenotic contacts the prootic, pterosphenoid and frontal and is pierced by a foramen conveying the otic branch of the facial to innervate the otic sensory canal. Internally the autosphenotic forms the anterior wall of the post-temporal fossa. Ridewood (1904) stated that in Elops saurus the anterior limit of the post-temporal fossa is formed by the pterosphenoid (


. Bulletin of the British Museum (Natural History) Geology Supplement. FOSSIL AND RECENT ors 15 I so. pto asp. Mot myp Fig. 4. Elops hawaiensis Regan. Neurocranium in orbital view. hyomandibular facet. The orbital face of the autosphenotic contacts the prootic, pterosphenoid and frontal and is pierced by a foramen conveying the otic branch of the facial to innervate the otic sensory canal. Internally the autosphenotic forms the anterior wall of the post-temporal fossa. Ridewood (1904) stated that in Elops saurus the anterior limit of the post-temporal fossa is formed by the pterosphenoid (alisphenoid). This is not the case in E. hawaiensis. The lateral margins of the heart-shaped optic foramen are formed by the ptero- sphenoid of either side which lies medial to both the prootic and autosphenotic. The surface of the pterosphenoid is marked by a groove which curves upwards and forwards from the opening to the pars jugularis. This groove contains the super- ficial ophthalmic branches of V and VII. The trochlear nerve passes out of the cranial cavity through the optic foramen. The orbitosphenoid lies anterior to the pterosphenoids. The anterior margin of the orbitosphenoid is perforated by a large foramen through which pass the olfactory tracts. Ventrally the orbitosphenoid is produced as a membranous interorbital septum. Posteriorly the orbitosphenoid forms the dorsal margin of the optic fora- men. The ventral margin of the optic foramen is formed by the ' wings ' of the ' Y '-shaped basisphenoid which contact the prootic and pterosphenoid of either side. The laterally flattened stem of the basisphenoid passes antero-ventrally to meet the parasphenoid so dividing the entrance to the posterior myodome. The basi- sphenoid stem ossifies late in ontogeny. The parasphenoid is long, extending from its contact with the vomer anteriorly to the hind edge of the basioccipital. Beneath the orbit the parasphenoid is relatively narrow but beneath the otic region it is broader


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