. Bulletin of the Museum of Comparative Zoology at Harvard College. Zoology. 164 BuUetin Museum of Comparative Zoology, Vol 142, No. 2 CO UJ o UJ a UJ OQ ID. 0- I- 2- 3- 4- 5- 6- 7- 8- 9-10-11-12-13-14-15-16-17-18-19-2021- SIZE CLASSES (EACH MM) cm. 24-25- 32- 51 Figure 8. Histogram of size distribution of lowland Carabidae in New Guinea. Each species is counted once, according tci its median lengtfi. Eacfi column represents number of species of which the mean length falls between mm; the actual numbers of species are given above the columns. The hatched portions of the first thr
. Bulletin of the Museum of Comparative Zoology at Harvard College. Zoology. 164 BuUetin Museum of Comparative Zoology, Vol 142, No. 2 CO UJ o UJ a UJ OQ ID. 0- I- 2- 3- 4- 5- 6- 7- 8- 9-10-11-12-13-14-15-16-17-18-19-2021- SIZE CLASSES (EACH MM) cm. 24-25- 32- 51 Figure 8. Histogram of size distribution of lowland Carabidae in New Guinea. Each species is counted once, according tci its median lengtfi. Eacfi column represents number of species of which the mean length falls between mm; the actual numbers of species are given above the columns. The hatched portions of the first three columns represent Tachys. See text for further explanation. may be one reason for their great success, and may enable them to Hve in the pres- ence of dominant ants more successfully than slightly larger Carabidae can do. Although several groups of Tachijs have certainly radiated to some extent in New Guinea, the number of species there is not due primarily to radiation but to the large number of separate stocks that have reached the island: my data sheets indi- cate at least 23 separate relationships be- tween different New Guincan Tachys and those of other areas, and this suggests at least 23 separate invasions of New Guinea by members of this one genus. In an attempt to get a more detailed explanation of the two size modes of low- land New Guinean Carabidae, I have made a size histogram (Fig. 9), of the 217 species that I found at Dobodura, indicat- ing the ecologic composition of the size classes. I have limited this diagram to species that I myself collected at one locality, because I know something ol their ecology. (See Ecology: habitats [24_ for further discussion of habitats of Ne^^' Guinean Carabidae.) This histogram sug gests that lowland mesophile Carabidae do have a bimodal size distribution in New Guinea. Mesophiles compete more di- rectly with ants than hydrophiles anc arboreal Carabidae do, and would be more likely to show the effects of competition. My tenta
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