. The cytoplasm of the plant cell. Plant cells and tissues; Protoplasm. Guilliermond - Atkinson 150 Cytoplasm other dyes which do not have a predilection for the vacuoles. More- over, the chondriosome-shaped vacuoles are of temporary nature and, even beginning with the first stages of cellular development, these vacuoles take in water and become transformed into large vacuoles, whereas it is known that the chondriosomes persist dur- ing the entire life of the cell. The only question that could be asked was whether, as we had supposed, the vacuoles were not derived in some cases from the chondr
. The cytoplasm of the plant cell. Plant cells and tissues; Protoplasm. Guilliermond - Atkinson 150 Cytoplasm other dyes which do not have a predilection for the vacuoles. More- over, the chondriosome-shaped vacuoles are of temporary nature and, even beginning with the first stages of cellular development, these vacuoles take in water and become transformed into large vacuoles, whereas it is known that the chondriosomes persist dur- ing the entire life of the cell. The only question that could be asked was whether, as we had supposed, the vacuoles were not derived in some cases from the chondriosomes. It was not logical to say that the chondriome and the vacuome were one and the same thing. The interpretation of Dangeard has therefore been abandoned for a long time now, even apparently by its author, although without explicit statement to that Fig. 94. — Barley root. Stages in development of the vacuolar system. 1-5, meristem; 6-8, adjacent region of differ- entiation; 9, mature cells of cortical parenchyma. Vital staining with neutral red. Our investigations, beginning with that period, gave more details and indeed confirmed the observations of P. A. Dangeard in so far as the development of the vacuoles is concerned, but they showed that the chondriosome-shaped vacuoles and the chon- driosomes have in common only their shape, and that the vacuome is a system completely independent of the chondriome. They showed, furthermore, that the vacuoles are not characterized by a specific substance corresponding to metachromatin and that the colloids which the vacuoles contain are substances of very diverse nature, having as a common property only their ability to absorb vital stains. These facts were afterwards verified by a large num- ber of cytologists and are today definitely accepted. The vacuoles do not, in general, stain by mitochondrial techniques and it is the very complex case of the leaflets of the rose which has caused all the difficulty. It has been se
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