Isador Michael Lerner (1910-1977, left), Russian geneticist and Ruggero Ceppellini (1917-1988, right), Italian geneticist. Lerner is best known for th

---

Isador Michael Lerner (1910-1977, left), Russian geneticist and Ruggero Ceppellini (1917-1988, right), Italian geneticist. Lerner is best known for the idea of genetic homoeostasis (also known as Lerner homeostasis) where populations maintain a steady genetic constitution over time. Ceppellini is best known for his work on the human leukocyte antigen (HLA). Photographed in San Francisco, USA, in 1954.

---

Size: 3604px × 2428px
Photo credit: © AMERICAN PHILOSOPHICAL SOCIETY/SCIENCE PHOTO LIBRARY / Alamy / Afripics
License: Licensed
Model Released: No

Keywords: 1900s, 1950s, 1954, 20th, adult, america, american, antigen, biological, biologist, biology, black--white, california, century, ceppellini, european, evolutionary, francisco, genetic, geneticist, geneticists, genetics, historical, history, hla, homeostasis, human, immunogenetics, immunological, immunology, isador, italian, lerner, leukocyte, male, man, men, michael, monochrome, north, outdoors, people, portrait, ruggero, russian, san, scientist, states, surname, united, usa