. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. New records of Vipera wagneri 227. Fig. 7: Habitat of V. wagneri, Kagizman area, NE Turkey. Photo Teynié. Morphologically, V. wagneri appears to be somewhat intermediate between V. xanthina and V. raddei in number of circumoculars (although still lacking a complete circumocular ring) and in number of ventrals. It agrees with V. bulgardaghica and V. bornmuelleri in low subcaudal counts. This character may be regarded primitive (Nilson & Andrén, 1986). If xanthina
. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. New records of Vipera wagneri 227. Fig. 7: Habitat of V. wagneri, Kagizman area, NE Turkey. Photo Teynié. Morphologically, V. wagneri appears to be somewhat intermediate between V. xanthina and V. raddei in number of circumoculars (although still lacking a complete circumocular ring) and in number of ventrals. It agrees with V. bulgardaghica and V. bornmuelleri in low subcaudal counts. This character may be regarded primitive (Nilson & Andrén, 1986). If xanthina and raddei are the most apomorphic species within the V. xanthina complex, then wagneri may be a relict form morphologically still close to the origin of xanthina. This makes also sense biogeographically, as the region south of the Cau- casus is likely to represent the evolutionary centre of this complex, from where the xanthina group spread to the west, whereas the raddei group radiated in eastward direction. It is noteworthy that the Munich specimen (o* 1) produced viable hybrids with a 9 V. raddei, but successful hybridizations between V. xanthina and V. raddei are also known. Acknowledgements We thank R. Gerstmeier, M. Kühbander, W. Schacht and K. Warncke for providing photo- graphs and locality data on the specimen they collected. Special thanks to U. Heckes for the drawing. Zusammenfassung Vipera wagneri Nilson & Andrén, 1984, bisher nur in einem 9 unsicherer Provenienz bekannt, wird anhand eines weiteren 9 und von 5 o* aus der Nordost-Türkei morphologisch charakteri- siert. Erstmals können Sexualdimorphismus und Lebendfärbung beschrieben werden. Der Artstatus für V. wagneri kann bestätigt werden. Die Art könnte als Reliktform gedeutet wer- den, die der Wurzel von V. xanthina nahe Please note that these images are extracted from scanned page images that may have been digitally enhanced for readability - coloration and appearance of these illustrati
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