. Bulletin of the British Museum (Natural History), Geology. FISHES From the Lebanon 273 has two epurals, not one as shown), and there are independent second and third uroneurals (un2, un3), as in Humbertia. There is a urodermal in Osmerns (Patterson ig68a : 230 ; Fig. 42) and Hypomesus (Fig. 43), but caudal scutes are absent (Weitz- man 1967b : 532). The neural spine of PU2 is short, as in Gaudryella and Humbertia, in all published illustrations of osmeroid tails (Gosline's individual of Hypomesus is abnormal in having two neural spines, one long and one short, on PU2 ; cf. Fig. 43). In Osmen


. Bulletin of the British Museum (Natural History), Geology. FISHES From the Lebanon 273 has two epurals, not one as shown), and there are independent second and third uroneurals (un2, un3), as in Humbertia. There is a urodermal in Osmerns (Patterson ig68a : 230 ; Fig. 42) and Hypomesus (Fig. 43), but caudal scutes are absent (Weitz- man 1967b : 532). The neural spine of PU2 is short, as in Gaudryella and Humbertia, in all published illustrations of osmeroid tails (Gosline's individual of Hypomesus is abnormal in having two neural spines, one long and one short, on PU2 ; cf. Fig. 43). In Osmenis, Berg illustrates short neural spines on both PU2 and PU3, a most unusual condition, while in the three dried skeletons of 0. eperlanus available to me, one (npu2, Fig. 42) has a short neural spine on PU2 and the other two have full neural spines. In general, osmeroids show a combination of primitive features (NPU2 short, three uroneurals, hypurals autogenous) and advanced features (no free U2, stegural fused with compound centrum and with no extension forward to PU2, two epurals, no caudal scutes) which distinguish them from both Gaudryella and Humbertia, but a Humbertia-\\ caudal skeleton could well be antecedent to the osmeroid Fig. 42. Osmerus eperlanus (L.), Recent. Caudal skeleton of a dried skeleton, BM(NH) unreg., 100 mm., Berlin. Arrows mark the outermost (unbranched) principal fin-rays, explanation of abbreviations p. 296. In stomiatoids, which Weitzman places as the closest relatives of osmeroids, the caudal skeleton is known in Polymetme, Vinciguerria (Weitzman 1967b, figs. 15, 10), Gonostoma, Maurolicus (Monod 1968, figs. 393, 394 ; Weitzman 1967a, fig. 31), Photichthys (Gosline i960, fig. 9) (all Gonostomatidae, the most primitive family according to Weitzman), Astronesthes, Borostomias (Weitzman 1967a, figs. 14, 31 ;. Please note that these images are extracted from scanned page images that may have been digitally enhanced for readability - coloratio


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