. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. Rana perezi in Tunisia 289 b c iiéÉiiiíiiiii 'IS'' "" ~ i -I 100 ms Fig. 2: Oscillograms of the calls of R. perezi. a: mating call; b: territorial call 1; c: transitional call; d: territorial call 2; e: territorial call 3; f: short high-frequency call (water temperature ° C in each case). nant frequency band is about 500 Hz wide and hes at 1991 ± 317 Hz (n = 110). It can be modulated in various ways; usually there is an upward modulation in the first c
. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. Rana perezi in Tunisia 289 b c iiéÉiiiíiiiii 'IS'' "" ~ i -I 100 ms Fig. 2: Oscillograms of the calls of R. perezi. a: mating call; b: territorial call 1; c: transitional call; d: territorial call 2; e: territorial call 3; f: short high-frequency call (water temperature ° C in each case). nant frequency band is about 500 Hz wide and hes at 1991 ± 317 Hz (n = 110). It can be modulated in various ways; usually there is an upward modulation in the first call of a series. The lower, usually somewhat weaker band Hes at 617 ± 115 Hz (n = 98) and may be un- modulated or may change in either the same direction as the upper band or the opposite direc- tion. There is a significant positive correlation between the level of the upper frequency band and the water temperature (Table 2). The lower band also appears to be positively correlated to some degree, but the correlation cannot be confirmed statistically with the available data (Table 2). 2. Territorial call 1 Territorial call 1 is the most common territorial call (Fig. 2). Its duration averages ms at 19° C and decreases significantly as the water temperature rises (Table 1). At 19° C this caU is composed of an average of pulses. The number of pulses decreases with increasing temperature (Table 1). Many territorial calls show some sign of subdivision into 2—6 pulse groups. However, unlike the mating call, these calls have no distinct intervals between the groups. Often a soft sound precedes the actual call. A sonagram of a territorial call 1 is shown in Fig. 4. Again there are two high-energy fre- quency bands, the upper and dominant one of which (1568 ± 306 Hz; n = 13) reaches significantly higher levels as the water temperature rises (Table 2). The lower frequency band is also positively correlated with the water temperature (Table 2). 3. Territorial call 2
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