. Bulletin of the Natural Histort Museum. Geology series. 122 RICHARDSON, RODRIGUEZ AND SUTHERLAND with narrow and coarse muri and an irregular reticulum are common accessory spores. Retusotriletes coronadus (Rodriguez, 1983: pi. 8, fig. 1) appears in the lower part of the zone. Remarks. The upper part of the EC Biozone (Aneurospora Sub- Biozone) is recognisable in England (Hereford and Worcester) in the Raglan Marl Formation below the first appearance of the MN Biozone in the St. Maughans Formation (Richardson, Rodriguez & Sutherland, 2000). In the La Vid section (Fig. 4)
. Bulletin of the Natural Histort Museum. Geology series. 122 RICHARDSON, RODRIGUEZ AND SUTHERLAND with narrow and coarse muri and an irregular reticulum are common accessory spores. Retusotriletes coronadus (Rodriguez, 1983: pi. 8, fig. 1) appears in the lower part of the zone. Remarks. The upper part of the EC Biozone (Aneurospora Sub- Biozone) is recognisable in England (Hereford and Worcester) in the Raglan Marl Formation below the first appearance of the MN Biozone in the St. Maughans Formation (Richardson, Rodriguez & Sutherland, 2000). In the La Vid section (Fig. 4) the earliest definite specimens of Scylaspora elegans and Aneurospora spp. occur together, consequently the lower part of the EC biozone has not been found there. Occurence. The zone is found in all four sections. Age. Upper Pffdoli to lowermost Devonian. The appearance of the genus Aneurospora is probably close to the base of the Devonian (Richardson, Ford & Parker 1984; Richardson, Rodriguez & Suther- land, 2000). Chitinozoans in the lower EC Biozone are of Middle Pffdoli age (basal EC, LaPeral,lowerEC,GerasandLa Vid sections). The same chitinozoans indicating a mid-Pfidoli age (Margachitina e/ega«.?jandPfidolitoearliestLochkovianage(t/n7or/2rm!0(/r«a)are both present in the upper part of the EC Biozone (Aneurospora Sub- Biozone) and in the top of the underlying H Biozone. Aneurospora spp. (A) Sub-Biozone (Figs 2-5) Reference section for base of sub-biozone. Upper San Pedro Formation, La Vid section, Cantabrian Mountains, NW Spain. The lowest occurrence of Aneurospora spp. was found in sample 10, c. 106 m above the base of the section. Definition. Appearance of the gtrwii, Aneurospora without proxi- mal papillae. Represents the interval between the appearance of Aneurospora and the incoming of Streelispora newportensis of the succeeding zone. Accessory species. Spores of the Chelinospora hemiesferica (Cramer & Diez) complex, C. cf. hemiesferica, C. cassicula Ri
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