. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. 66 Bonner zoologische Beiträge 53 (2004) certain monophyly of genera as Calicnemia and Coelic- cia, which partly have been defined on the base of vari- able (polymorphic) wing venational traits of which the group-defining properties very much depend on our definition of the character states concerned. Secondly, it is necessary to newly define the generic borders be- tween Coeliccia and Asthenocnemis. Several new as. Fig. 53: Map of Malesia, with distribution range o
. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. 66 Bonner zoologische Beiträge 53 (2004) certain monophyly of genera as Calicnemia and Coelic- cia, which partly have been defined on the base of vari- able (polymorphic) wing venational traits of which the group-defining properties very much depend on our definition of the character states concerned. Secondly, it is necessary to newly define the generic borders be- tween Coeliccia and Asthenocnemis. Several new as. Fig. 53: Map of Malesia, with distribution range of Calicnemiinae indicated. well as described species should be transferred to the latter genus; distinct putative synapomorphies have been found (Dukstra & Gassmann, in prep.). Thirdly, by increasing the range of taxa and of both morphological ( endosceletal) and molecular characters, the am- bigious phylogenetic position of Asthenocnemis could possibly be resolved. Implications for biogeography As a group of insects occuring to the west and to the east of the Wallace line (Fig. 53), the Calicnemiinae are of special interest for biogeographical studies in the Indo-Australian transition zone. The phylogenetic hy- pothesis presented above allows for the postulation of active or passive dispersal events both into the East as well as, subsequently, back into the West ( the Phil- ippines). Since a detailed scenario for the distributional history of the group has been developed elsewhere in the context of the historical biogeography of the fresh- water biota in Southeast Asia (Van Tol & GASSMANN, submitted), the reader is referred to that forthcoming paper. Acknowledgments. It is an honour to me to dedicate the present paper to the memory of Professor Cías M. Naumann (Zoological Research Institute and Museum Alexander Koenig Bonn, and University of Bonn/ Germany), whom I owe a long lasting motivation and en- thusiasm to work in the field of organismic biology;
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