. Bulletin of the Natural Histort Museum. Geology series. 104 SMITH ETAL. less uniserial to peristome, with no pore crowding whatsoever. Small buccal pores are present at the peristomial margin. Periproct oval, longitudinal, in distinct anal sulcus opening closer to the posterior margin than the apical system. Occurrence. Maastrichtian, horizon 2 (top) and 4, Santander; Maastrichtian, horizon 6, Olazagutia Pass, Navarra province. This species also occurs in the upper Lower to lower Upper Maastrichtian of the Maastricht district. The Netherlands and Belgium (Van der Ham et al., 1987), the
. Bulletin of the Natural Histort Museum. Geology series. 104 SMITH ETAL. less uniserial to peristome, with no pore crowding whatsoever. Small buccal pores are present at the peristomial margin. Periproct oval, longitudinal, in distinct anal sulcus opening closer to the posterior margin than the apical system. Occurrence. Maastrichtian, horizon 2 (top) and 4, Santander; Maastrichtian, horizon 6, Olazagutia Pass, Navarra province. This species also occurs in the upper Lower to lower Upper Maastrichtian of the Maastricht district. The Netherlands and Belgium (Van der Ham et al., 1987), the Maastrichtian 'Craie phosphatee" at Ciply, Belgium (Lambert, 1898b) and the Maastrichtian at Fresville, Manche, France (Lambert collection). Material studied. BMNH EE4403, EE6148-49, MGB 37519. Remarks. This is a small species of juvenile appearance (simple short petals, simple phyllodes, absence of bourrelets). There are several names available for such forms, but they lack distinctive characters and are virtually impossible to separate. Consequently, we group all together here under the name A^. coravium Defrance. This species is readily distinguished from the co-occurring A'. scrobiculatus by its shape, which is much less tumid and with a deeply sunken oral surface, and by its phyllodes, which are com- pletely undifferentiated. It differs from A', geayi in having a more posterior periproctal opening. Nucleopygus scrobiculatus (Goldfuss, 1829) PI. 4, figs 1-6; Fig. 12 1829 Nucleolites scrobiculatus Goldfuss: 138, pi. 43, fig. 3. 1935 Lvchnidius scrobiculatus (Goldfuss); Smiser: 49, pi. 4, fig. 10. 1957 Nucleopygus scrobiculatus (Goldfuss); Engel & Meijer: 88, pi. L 1987 Nucleopygus scrobiculatus (Goldfuss); Van der Ham et al.: 30, pi. 11, fig. 3. Diagnosis. Test up to 26 mm in length, ovate; width 85-90% of length, widest centrally; height 60-65% of length, tallest behind apical disc. Apical disc a little anterior of centre (40% of length); tetrabasal with four gonop
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