. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. 348 Bonner zoologische Beiträge 53 (2004). Fig. 11: Agathon setosus sp. n. a, pupa in dorsal view; b, front end seen from left side; c, left respiratory organ, lamella 4 partly cut to expose lamellae two and 3. Figures a and b to same scale. Genus Philorus Kellogg, 1903 Philorus Kellogg, 1903 (Kellogg 1903: 199). Type species: Blepharocera yosemite (Osten-Sacken, 1877), by subsequent designation of Coquillett, 1910 (CO- quillett 1910: 588). The phylogenetic rel
. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. 348 Bonner zoologische Beiträge 53 (2004). Fig. 11: Agathon setosus sp. n. a, pupa in dorsal view; b, front end seen from left side; c, left respiratory organ, lamella 4 partly cut to expose lamellae two and 3. Figures a and b to same scale. Genus Philorus Kellogg, 1903 Philorus Kellogg, 1903 (Kellogg 1903: 199). Type species: Blepharocera yosemite (Osten-Sacken, 1877), by subsequent designation of Coquillett, 1910 (CO- quillett 1910: 588). The phylogenetic relations of Philorus among Ble- pharicerini are doubtful because the genus still shares presumably primitive characters with Bibiocephala and Agathon; see under Bibiocephala. At the same time, the radial vein is more reduced (two short branches on a long stalk, instead of long independent branches) than in other Blepharicerini, supporting monophyly of genus Philorus. The pupal respiratory organ of four low, widely divergent lamellae is another obvious apomor- phy. The genus comprises almost 25 species from mainland Asia (south to Assam and west to Tian Shan), Japan, and western North America. Grouping of included spe- cies is not yet generally clear. However, the new Rus- sian species clearly belongs to a monophyletic group distinguished by: lateral digitiform appendages on male tergite 9; male with extremely short ventral bridge from which the bag-like, uncrested vesica extends freely for- ward; pupal respiratory organs medially touching, la- mellae very low, crest-like, divergent. The name Euliponeura Tonnoir, 1930 would be avail- able for this group which comprises P. asiaticus Brodsky, 1972, P. assamensis (Tonnoir, 1930), P. cho- senensis Kitakami, 1931, P. horai (Tonnoir, 1930), P. longirostris Kitakami, 1931, P. minor Kitakami, 1931; not all of the three characters listed above are known for each included species. Recognising Euliponeura would turn the genus Philorus para
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