. Bulletin of the British Museum (Natural History) Entomology. SYSTEMATICS OF THE PLUSIINAE 121 The 647-step cladograms Following removal of the outgroups, further global branch-swapping with the multiple parsimony option discovered a set of 16 cladograms. After reinstatement of the outgroups, these cladograms were found to be 647 steps long, one step shorter than those found in the earlier analyses that were described above. The 647-step cladograms are of similar topology to the rejected 648-step subcladogram 12, in that Rachiplusia forms the sister-group of the Argyrogrammatini, but differ i
. Bulletin of the British Museum (Natural History) Entomology. SYSTEMATICS OF THE PLUSIINAE 121 The 647-step cladograms Following removal of the outgroups, further global branch-swapping with the multiple parsimony option discovered a set of 16 cladograms. After reinstatement of the outgroups, these cladograms were found to be 647 steps long, one step shorter than those found in the earlier analyses that were described above. The 647-step cladograms are of similar topology to the rejected 648-step subcladogram 12, in that Rachiplusia forms the sister-group of the Argyrogrammatini, but differ in the branching sequence of the subtribes of the Plusiini (Fig. 26). Whereas, on subcladogram 12 of the 648-step cladograms, the Autoplusiina (less Rachiplusia) form the sister-group of the Argyrogrammatini + Rachiplusia, on the 647-step cladograms, it is the Plusiina that occupy this position. The 16 alternatives arise from the mutually exclusive topologies referred to previously as subcladograms 1/2 (Agraphai'Plusiopalpa), 3/4 (Plusidia) and 5/6/7/8 (Auto- graphalComutiplusialPlusia). The presence of further alternatives in the outgroup taxa was not observed but all of the topologies found previously would be expected. Reversing the order of branching of the Plusiina and Autoplusiina (less Rachiplusia) appears to preclude the alternative topology that places Loboplusia as the sister-group to Erythroplasia. ARGYROGRAMMATINI Rachiplusia PLUSIINA AUTOPLUSIINA EUCHALCIINA ABROSTOLINI Fig. 26 The branching sequence of the tribes and subtribes on the 647-step cladograms. Note that the position of Rachiplusia is the same as on subcladogram 12 of the 648-step cladograms (Fig. 19) but that the Autoplusiina and the Plusiina have exchanged positions. However, the 647-step cladograms (which, given that the branch-swapping procedure ran to comple- tion, are most likely the shortest for this data set) are based upon the same characters (with one exception) as the previously rejected 648-st
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