. Handbook of flower pollination : based upon Hermann Mu?ller's work 'The fertilisation of flowers by insects' . Fertilization of plants. THYMELAEACEAE 357 2503. D. Mezereum L. (Herm. Miiller, 'Weit. Beob.,' II, p. 236, 'Alpen- blumen/ p. 207 ; Schulz, 'Beitrage,' II, pp. 159-60; Kerner, 'Nat. Hist. PI.,' Eng. Ed. I, II, p. 3CI ; Kirchner, 'Flora v. Stuttgart,' p. 423 ; Ludwig, ' Adynamandrie V. Erodium macradenum u. s. w.'; Knuth, ' Bloemenbiol. Bijdragen.'—The bright purple flowers of this species are strongly fragrant, thus attracting numerous bees, flies, and Lepidoptera, especially as the


. Handbook of flower pollination : based upon Hermann Mu?ller's work 'The fertilisation of flowers by insects' . Fertilization of plants. THYMELAEACEAE 357 2503. D. Mezereum L. (Herm. Miiller, 'Weit. Beob.,' II, p. 236, 'Alpen- blumen/ p. 207 ; Schulz, 'Beitrage,' II, pp. 159-60; Kerner, 'Nat. Hist. PI.,' Eng. Ed. I, II, p. 3CI ; Kirchner, 'Flora v. Stuttgart,' p. 423 ; Ludwig, ' Adynamandrie V. Erodium macradenum u. s. w.'; Knuth, ' Bloemenbiol. Bijdragen.'—The bright purple flowers of this species are strongly fragrant, thus attracting numerous bees, flies, and Lepidoptera, especially as the crowded blossoms are not hidden by foliage- leaves ; nectar is secreted by the base of the ovary. Hermann Miiller says that the proboscis of an insect probing for this first brushes against the two rows of anthers in the corolla-tube without removing their pollen, and then touches the stigma below them. Pollination will be eff'ected if another flower has been previously visited. When the proboscis, wetted with nectar, is withdrawn, pollen adheres to it and is transferred to flowers visited later. If insect-visits fail, automatic self-pollination takes place by fall of pollen; this, however, does not seem to be always effective, for all flowers do not set fruits. Kerner says that autogamy seldom occurs on account of the horizontal position of the flowers. Schulz observed gynomonoecism and, rarely, gynodioecism; he also found the flowers at Halle self-fertile. Ludwig, on the contrary, observed adynamandry at Greiz. He transplanted two stocks from different parts of a wood into his garden. They set abundant fruits for eight years, and then one of them died. In spite of frequent insect- visits and artificial transfer of the normal pollen to the fully mature stigmas, no fruits were set by the other plant. Ludwig adds :— ' This case of adynamandry deserves special notice, as A. Schulz fig. 358. Daphne Mezereum, L. (after Herm. Muller). found the species to be self-fertile f- a f


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