. Bulletin of the British Museum (Natural History), Geology. 288 THE EARLIEST GOATS AND OTHER ANTELOPES post cornual groove, orbital rims strongly projecting, braincase not strongly angled, longer braincase, frontals low between the horn bases, no raising of the mid-frontals suture, infraorbital foramen high over P3, a less strong median ridge on the occiput, poor styles and ribs on the upper molars, the lateral wall of P4 indented in front of the hypoconid, P3 relatively smaller, and the front and back edges of P3 and P4 not set transversely. Phytogeny of spiral horned A ntilopini On the ques
. Bulletin of the British Museum (Natural History), Geology. 288 THE EARLIEST GOATS AND OTHER ANTELOPES post cornual groove, orbital rims strongly projecting, braincase not strongly angled, longer braincase, frontals low between the horn bases, no raising of the mid-frontals suture, infraorbital foramen high over P3, a less strong median ridge on the occiput, poor styles and ribs on the upper molars, the lateral wall of P4 indented in front of the hypoconid, P3 relatively smaller, and the front and back edges of P3 and P4 not set transversely. Phytogeny of spiral horned A ntilopini On the question of phylogeny, not a lot can be said, particularly in the absence of detailed time correlations of the sites where the varieties of horn core types occur. It is possible to conceive that the Maragha and Samos forms of P. rotundicornis are later than that in Pikermi, in so far as they are more gracile and therefore remote from P. houtumschindleri. However this conclusion is the more doubtful by the fact that the Pikermi form of Sporadotragus appears more advanced than at Samos. Bearing in mind the possibility of a time span at Samos, it is as well to follow the speculation no further. On the whole P. rotundicornis shows more signs of ancestry to A. cervicapra than any other of the Pliocene species, and it is a pity that it is less completely known than P. houtumschindleri or Protragelaphus Fig. 16. Possible relationships of bovids mentioned in this paper. Names of species and genera which have been dealt with at length are underlined. The middle horizontal band contains forms known from Samos and other sites of broadly the same age. The upper band is for later forms, and the lower one is for a pre-Samos time level. Precise origins are not shown for Spirocerus, Ovibos and Please note that these images are extracted from scanned page images that may have been digitally enhanced for readability - coloration and appearance of these illustrations may not
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