. The Canadian field-naturalist. 1979 Roberts and Lewin: Amphibian Populations, Alberta 149 55 50H 45 40H « uj 35 OD = 301 z < m =? 25H 20- lO 5- KEY HABITAT types: 1 Water Wet 2 Sedge Fen 3 Grass Meadow 4 Willow-bog 5 Aspen Poplar 6 Black Spruce 7 White Spruce 8 Upland Mixed Wood 9 Jack Pine Dry symbols; WOOD FROG Adults (N=I5) ^ Yearlings(N = IO) Young-of-year(N=5) rlflirK. i^r:? I 2 3 4 5 6 7 8 9 MAY 23456789 JU NE 23456789 123456789 JULY AUGUST habitat types Figure 2. Habitat utilization by Wood Frogs in northeastern Alberta during May-August 1977. grass meadows, willow bogs, and aspen
. The Canadian field-naturalist. 1979 Roberts and Lewin: Amphibian Populations, Alberta 149 55 50H 45 40H « uj 35 OD = 301 z < m =? 25H 20- lO 5- KEY HABITAT types: 1 Water Wet 2 Sedge Fen 3 Grass Meadow 4 Willow-bog 5 Aspen Poplar 6 Black Spruce 7 White Spruce 8 Upland Mixed Wood 9 Jack Pine Dry symbols; WOOD FROG Adults (N=I5) ^ Yearlings(N = IO) Young-of-year(N=5) rlflirK. i^r:? I 2 3 4 5 6 7 8 9 MAY 23456789 JU NE 23456789 123456789 JULY AUGUST habitat types Figure 2. Habitat utilization by Wood Frogs in northeastern Alberta during May-August 1977. grass meadows, willow bogs, and aspen poplar habitat, which are the most moist terrestrial habitats available. Heatewole (1961) regards the Wood Frog as a forest-dwelling species and suggests that the shade provided by trees is an important factor in determining habitat selec- tion. He demonstrated experimentally that substrate structure and substrate moisture are important factors governing the selection of microhabitat. Bellis (1962) states that Wood Frogs are restricted to forested areas, particu- larly lowland bogs. Shelford (1913) demon- strated that Wood Frogs prefer an environment with moist air. Marshall and Buell (1955) found that in exposed areas with sparse Wood Frog populations the vapor pressure deficit was higher than in shaded tamarack {Larix larcinia) habitat with a dense Wood Frog population. It thus appears that moisture content of both the substrate and air are important factors in determining the suitability of habitat types for Wood Frogs. Tall grasses, willows, and aspen poplars growing in moist soil in low-lying areas provide a favorable environment for Wood Frogs within the study area, while much of the forest habitat, especially Jack Pine area, is too dry with respect to substrate moisture and perhaps the humidity of the air. Heatwole (1961) suggests that the Wood Frog is less terrestrial than previously thought (see Wright and Wright 1949) and points out that its association with woods an
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