. California fish and game. Fisheries -- California; Game and game-birds -- California; Fishes -- California; Animal Population Groups; Pêches; Gibier; Poissons. 176 CALIFORNIA FISH AND CAME nia. Lee et al. (1980) listed the four subspecies studied here, but described them as poorly defined with the validity of all subspecies questionable. Moyle (1976) recognized the four subspecies studied here, but cited the need for a thorough systematic review. Lake. â Sacramento sucker â¢Goose Lake sucker DHumboldt sucker OPajaro sucker FIGURE 1. Map of northern California and southwestern Oregon showing


. California fish and game. Fisheries -- California; Game and game-birds -- California; Fishes -- California; Animal Population Groups; Pêches; Gibier; Poissons. 176 CALIFORNIA FISH AND CAME nia. Lee et al. (1980) listed the four subspecies studied here, but described them as poorly defined with the validity of all subspecies questionable. Moyle (1976) recognized the four subspecies studied here, but cited the need for a thorough systematic review. Lake. â Sacramento sucker â¢Goose Lake sucker DHumboldt sucker OPajaro sucker FIGURE 1. Map of northern California and southwestern Oregon showing distributions of Catostomus occidentalis. The river or lake system of each population studied is labeled. The very use of the term subspecies as an evolutionary lineage has also been confused and controversial. A subspecies has been defined as "an aggregate of phenotypically similar populations of a species, inhabiting a geographic subdi- vision of the range of a species, and differing taxonomically from other popu- lations of the species" (Mayr 1969). The degree of taxonomical difference used to delineate subspecies has varied. Wilson and Brown (1953) called for the aban- donment of the subspecies trinomial and its replacement by a system of refer- ence based on relevant geographical names. This study involved analysis of adult and juvenile characters to identify and compare subspecies. Emphasis was placed on comparison of meristic characters because morphological features of suckers may differ between sexes. Cluster analysis, scale count comparisons and discriminant analysis were used to exam- ine relationships within C. occidentalis. Relatively few taxonomic studies involv- ing catostomids have used multivariate statistical techniques. Ferris and Whitt (1978) used cluster analysis based on phenetic similarities to determine rela- tionships between 30 species of catostomids. Smith and Koehn (1971) did phenetic and cladistic studies on 16 species of catostomids


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