. British journal of entomology and natural history. Natural history; Entomology. BR. J. ENT. NAT. HIST., 9: 1996 WING HOMOEOSIS IN PARARGE AEGERL^f^^^^f^ (LEPIDOPTERA: SATYRIDAE) Leonard Winokur \ ' c 1 1997 ) 55 Palmer Park Avenue, Reading, Berkshire RG6 1DP\^ I ie_. _« A female speckled wood butterfly, Pararge aegeria L. subspecies oblita Harrison, emerged 1995 among a bred stock derived from two females collected at Glen Lonan, east of Oban, Argyll, (56° 24' N, 5 22' W). Pattern typical of left hindwing underside spaces 2 and 3 (s2 and s3) was reproduced at correspondi


. British journal of entomology and natural history. Natural history; Entomology. BR. J. ENT. NAT. HIST., 9: 1996 WING HOMOEOSIS IN PARARGE AEGERL^f^^^^f^ (LEPIDOPTERA: SATYRIDAE) Leonard Winokur \ ' c 1 1997 ) 55 Palmer Park Avenue, Reading, Berkshire RG6 1DP\^ I ie_. _« A female speckled wood butterfly, Pararge aegeria L. subspecies oblita Harrison, emerged 1995 among a bred stock derived from two females collected at Glen Lonan, east of Oban, Argyll, (56° 24' N, 5 22' W). Pattern typical of left hindwing underside spaces 2 and 3 (s2 and s3) was reproduced at corresponding positions in left forewing underside s2 and s3 (Fig. 1). There was no corresponding pattern transformation on the right forewing underside nor on the upper surface. Homoeosis refers to the development of tissue typical of one part of an organism at a position typical of another tissue type. In Lepidoptera this is typically recognized by an area on one wing showing the pattern of another wing. Homoeosis is rare in Lepidoptera (Ford, 1957) but has been reported in several families including the Satyridae. Insect forewings belong to the middle thoracic segment or mesothorax, and the hindwings to the hind thoracic segment or metathorax. The incidence of different types of homoeotic transformation in Lepidoptera, has revealed also that each wing is divided further into anterior and posterior developmental units or 'compartments', along a baso-distal axis that borders s5 and s6 (Sibatani, 1980). In the following discussion, si to s5 on the forewing and si to s4 on the hindwing represent the 'posterior' of the respective wing, while the forewing area between s6 and the costa and the hindwing area between s5 and the costa represent the 'anterior' of the respective wing. The most common type of homoeosis in Satyridae concerns the presence within the underside posterior hindwing of the underside posterior forewing pattern. The present case is unusual therefore in demonstrating the p


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