. Bulletin of the Museum of Comparative Zoology at Harvard College. Zoology. k-AND-^MT EN PLD. PRD HYD Figure 26. Tooth of Villarroelomys bolivianus Hartenberger compared with dm4 of Cephalomys arcidens, x5. A. GB no. 014 redrawn from Hartenberger, 1975: Fig. 2a, and restored according to our interpretation of its probable structure. B. Rdm4 of Ceptialomys arcidens, ACM no. 3013, redrawn and reversed from Wood and Patterson, 1959: Fig. 20A. Abbreviations: AND—area derived from anteroconid; EN—entoconid; HYD—hypoconid; MT- lophid; PRD—protoconid. -metaconid; PLD—postero- how this tooth could ha


. Bulletin of the Museum of Comparative Zoology at Harvard College. Zoology. k-AND-^MT EN PLD. PRD HYD Figure 26. Tooth of Villarroelomys bolivianus Hartenberger compared with dm4 of Cephalomys arcidens, x5. A. GB no. 014 redrawn from Hartenberger, 1975: Fig. 2a, and restored according to our interpretation of its probable structure. B. Rdm4 of Ceptialomys arcidens, ACM no. 3013, redrawn and reversed from Wood and Patterson, 1959: Fig. 20A. Abbreviations: AND—area derived from anteroconid; EN—entoconid; HYD—hypoconid; MT- lophid; PRD—protoconid. -metaconid; PLD—postero- how this tooth could have erupted except after dm^, which suggests that it might have been a P^ that was shed along with dm^ (Wood, 1969; 1970b: 245-246). One badly broken dm4 is present in the Princeton collection (PU no. 21987, Fig. 24G). This preserves no more than the posterior half of the pattern, which is the most molarifomi part of the caviomorph dnii. In view of this, little more can be said of the specimen. Lavocat (1976: 39; PL 2, Fig. 5) described a similarly broken and more worn specimen in a ramus frag- ment that also includes Mj. Lavocat (1976: 40-41) and Hoffstetter (1976: 10, n.) concluded that the type of Villarroelomys bolivianus (Hartenber- ger, 1975) was a dm4, not an Mg, and sus- pected that it may have been a young in- dividual oi Branisamys, in which Mj had not yet erupted. Lavocat further stated that Hartenberger was in essential agree- ment with him on this point (Lavocat, 1976: 40). We had independently arrived at the same conclusion. Certainly this tooth has nothing to do with the hydro- choerids. Because Lavocat felt that there was some possible uncertainty as to the reference to Branisamys, since the tooth of Villarroelomys bolivianus might rep- resent an otherwise unknown genus closely related to Branisamys, he be- lieved that the name Villarroelomys should be retained for the time being. Hartenberger's illustration of the type oi Villarroelomys bolivianus (197


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