. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. Geographie variation in Gallotia galloti 537 The 'dispersal' interpretation of the phylogenetic tree does not indicate an outright secondary origin of the G. eisentrauti ( Tenerife), G galloti (S. Tene- rife) contact within Tenerife as it possibly could have. That is, it is possible to envisage an origin in La Palma, colonization of Tenerife from La Palma, colonization of Hierro from La Palma, Gomera from Hierro and S. Tenerife from Gomera. This would lead


. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. Geographie variation in Gallotia galloti 537 The 'dispersal' interpretation of the phylogenetic tree does not indicate an outright secondary origin of the G. eisentrauti ( Tenerife), G galloti (S. Tene- rife) contact within Tenerife as it possibly could have. That is, it is possible to envisage an origin in La Palma, colonization of Tenerife from La Palma, colonization of Hierro from La Palma, Gomera from Hierro and S. Tenerife from Gomera. This would lead to a secondary contact between G galloti gal- loti and G. galloti eisentrauti. However, both hypothesized pathways (Fig. 2b, 3 b) are consistent with a primary origin of this zone in situ. It would be unwise to preclude a secondary origin from this zone within Tenerife on the basis of the preliminary analysis particularly since one can envisage a temporary vica- riance of northern and southern populations within Tenerife (resulting in a secon- dary contact) that would be consistent with the colonization sequence of Figs. 2b and 3 b. The pattern of dispersal indicated by the out-group root (Fig. 2 b) conforms to the past and current distribution of the species, G simonyi. Gallotia simonyi has a relict distribution which may typify a late stage in the taxon cycle (Wilson 1961) where an early species has differentiated but has been out-competed by newer species and has consequently become extinct over much of its former range. In the west, G. simonyi has a small relict population on one cliff in Hierro and was until recently found on the small islets off Hierro and Tenerife (Baez & Bravo 1983). However, fossil evidence indicates that it previously had a wider western distribution across Hierro, Gomera and Tenerife (but not La Palma). The absence of G. simonyi on La Palma would allow its colonization by a popu- lation that evolved into G. galloti as suggested by the La Palma o


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