. Bulletin of the Museum of Comparative Zoology at Harvard College. Zoology. C E L. Figure 20. Range of additional trees (of length 27 or less) derivable from the character data entered into the analysis. 1A (length 24), 2A-F (length 26), and 3A-B (length 27). C = Cichlidae, P = Pomacentridae, E = Embiotocidae and L = Labridae. the predominance of the original com- plex may be expected to be reduced. This isn't, of course, to say that those original features suddenly become un- important or insignificant, but only that they no longer predominate. In view of our conscious effort to lo- cate fea


. Bulletin of the Museum of Comparative Zoology at Harvard College. Zoology. C E L. Figure 20. Range of additional trees (of length 27 or less) derivable from the character data entered into the analysis. 1A (length 24), 2A-F (length 26), and 3A-B (length 27). C = Cichlidae, P = Pomacentridae, E = Embiotocidae and L = Labridae. the predominance of the original com- plex may be expected to be reduced. This isn't, of course, to say that those original features suddenly become un- important or insignificant, but only that they no longer predominate. In view of our conscious effort to lo- cate features other than pharyngeal ones uniting the Labroidei and our in- ability to find any, we suggest that it is improbable that investigator bias is re- sponsible for the importance of the pharynx in diagnosing the clade. Of course we cannot rule out the possibil- ity that other morphological informa- tion does exist and that we have simply not found it yet, but our hypothesis is that such data do not exist. 4. Predominance of the region may sim- ply be a reflection of structural (and/ or functional) complexity. As Lauder (1981) quite correctly pointed out, few morphologists have explicitly consid- ered the influence of complexity upon patterns of morphological change. In- tuitively at least, it seems that complex systems have a higher likelihood of change than simple ones. If complexity is defined as the number of parameters needed to describe form (Lauder, 1981; Vermeij, 1973), then an increase of complexity will automatically increase the number of possibilities for change in the component elements and in their relations to one another. Complex sys- tems have more potentially stable in- termediate states and have, therefore, options for change in design at each level (Lauder, 1981; Simon, 1962). The euteleostean pharynx is a highly complex construction, composed of many elements and numerous struc-. Please note that these images are extracted from scanned page images that may


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