. Bonner zoologische Monographien. Zoology. HUBER. REVISION AND CLADISTIC ANALYSIS Ol- PI/()/x:i'S AND CLOSl l Y Ria Ariari AXA (ARANEAE, l'HOLCIDAE). FIG. 1707, 1708. Known distributions of Pholciis bessus and of Ph. manueli in Asia (1707) and North America (1708). Pholcus crypticolens Bösenberg & Strand, 1906 Figs. 1651-1653, 1669, 1670, 1709-1721, 1722- 1724 Pholcus crypticolens Bösenberg & Strand 1906: 127, pl. 6, flg. 72, pl. 15, figs. 440a-f Saito 1934: 299 (not seen). Uyemura 1937: III, figs. A-F (not seen). Saito 1959: 65, figs. 51a-e. Yaginuma 1960: 48, fig. 96; 1970: 646; 198
. Bonner zoologische Monographien. Zoology. HUBER. REVISION AND CLADISTIC ANALYSIS Ol- PI/()/x:i'S AND CLOSl l Y Ria Ariari AXA (ARANEAE, l'HOLCIDAE). FIG. 1707, 1708. Known distributions of Pholciis bessus and of Ph. manueli in Asia (1707) and North America (1708). Pholcus crypticolens Bösenberg & Strand, 1906 Figs. 1651-1653, 1669, 1670, 1709-1721, 1722- 1724 Pholcus crypticolens Bösenberg & Strand 1906: 127, pl. 6, flg. 72, pl. 15, figs. 440a-f Saito 1934: 299 (not seen). Uyemura 1937: III, figs. A-F (not seen). Saito 1959: 65, figs. 51a-e. Yaginuma 1960: 48, fig. 96; 1970: 646; 1986: 29, figs. 2e, p, pl. 6, fig. 10. Chikuni 1989: 28, fig. 3. Misidentifications: see Ph. zichyi and Ph. spilis below. ''Pholcus zichyf (misidentification): Irie 2009: 110, figs. (2-2-13) 33-35. Note. Mainland (Russian, Chinese, Korean) speci- mens have traditionally been identified either as Ph. crypticolens or Ph. minutus (for references see Ph. zichyi below). Marusik & Koponen (2000) synony- mized Ph. crypticolens and Ph. minutus with Ph. zichyi, but the senior synonym Ph. zichyi was sup- pressed by Platnick (2001) for lack of usage. I have compared the type series of Ph. minutus with Chinese specimens from near the type locality of Ph. zichyi and found them to be identical, in agreement with Marusik & Koponen (2000). However, Japanese specimens are consistently different from mainland specimens, so Ph. crypticolens and Ph. zichyi are both considered valid species. Types. Two female and 5 juvenile syntypes from Ja- pan, near Nagasaki [-32°54'N, 129°54'E], 1882 (Dönitz), in SMF (2943), examined. Diagnosis. Easily distinguished from most congeners by distinctive color pattern of carapace and abdomen (dorsally and ventrally) (Figs. 1651, 1653) and of legs (sprinkled with light brown spots); also by morphol- ogy of male palp (Figs. 1709, 1710; shapes of pro- cursus, uncus, and appendix) and female genitalia (Figs. 1669, 1712); from the very similar Ph. zichyi and Ph
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