. Bulletin of the British Museum (Natural History) Zoology. PHARYNGOBRANCHIAL ORGAN OF MUGILID FISHES 129 regarded Mugil labiosus Val. as a species of Oedalechilus which Schultz (1953) had made the type species of Plicomugil. There are, however, distinct differences in the PBO's, and in the nature of the lip folding, tooth shape and lower jaw shape between O. labeo and O. labiosus (p. 125). We thus accept Plicomugil as a valid genus to contain O. labiosus which exhibits the expanded denticulate area typical of the derived condition (node 5 on Fig. 20), (cf. O. labeo). Mugil has a variable PBO
. Bulletin of the British Museum (Natural History) Zoology. PHARYNGOBRANCHIAL ORGAN OF MUGILID FISHES 129 regarded Mugil labiosus Val. as a species of Oedalechilus which Schultz (1953) had made the type species of Plicomugil. There are, however, distinct differences in the PBO's, and in the nature of the lip folding, tooth shape and lower jaw shape between O. labeo and O. labiosus (p. 125). We thus accept Plicomugil as a valid genus to contain O. labiosus which exhibits the expanded denticulate area typical of the derived condition (node 5 on Fig. 20), (cf. O. labeo). Mugil has a variable PBO morphology and among the species examined, five groups can be distinguished (see p. 125 for description of the features delimiting these groups), viz: 1) including M. cephalus (type species), bananensis, curema, curvidens, incilis and trichodon; 2) M. capurii, considered by Ingham (1952) the most primitive Mugil species (certainly the wide and shallow separation of the denticulate cushions and large proximal teeth are plesiomorphic features); 3) M. liza which more closely resembles Liza species; 4) M. thoburni and M. setosus, which are certainly closely related species and may represent the respective ends of a clinal distribution, the former in the Galapagos, the latter at the western coast of Mexico (Schultz (1946) recognised Xenomugil to contain M. thorburni); 5) M. hospes, which in the feature of its vertical sulcus most closely resembles Neomyxus leuciscus, but the overall PBO morphology and valve size suggests that it is a specialised variant of the M. cephalus group. Now considering the advanced genera which exhibit the more plesiomorphic condition of a relatively small denticulate area (ie., terminal group of Fig. 20), Sicamugil has a Liza-like PBO morphology although with specialised features (p. 124). Schultz (1946) synonymised Sicamugil with Tr achy stoma {—. Please note that these images are extracted from scanned page images that may have been digitally enhanced
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