. The Ecology of arboreal folivores : a symposium held at the Conservation and Research Center, National Zoological Park, Smithsonian Institution, May 29-31, 1975 . Figure 9. Distribution of locomotor patterns on different-sized supports in locomotion during travel and locomotion during feed- ing. Horizontal axis shows the contribution of different patterns in each kilometer of progression; vertical axis gives the proportion of that distance that is moved on different types of supports. In P. melalophos hopping is separated from other forms of leaping. For leaping, take-off supports and landin


. The Ecology of arboreal folivores : a symposium held at the Conservation and Research Center, National Zoological Park, Smithsonian Institution, May 29-31, 1975 . Figure 9. Distribution of locomotor patterns on different-sized supports in locomotion during travel and locomotion during feed- ing. Horizontal axis shows the contribution of different patterns in each kilometer of progression; vertical axis gives the proportion of that distance that is moved on different types of supports. In P. melalophos hopping is separated from other forms of leaping. For leaping, take-off supports and landing supports are shown sep- arately. quadrupedal walking and running or leaping and hopping; climbing and armswinging bouts are shorter. Quadrupedal bouts of P. obscura are signifi- cantly longer than bouts of any other locomotor activity of either monkey. This reflects the extensive utilization of continuous main canopy supports by that species. The leaps of P. obscura are shorter than those of P. melalophos. In both species, the relative use of locomotor pat- terns during feeding is significantly different from that during travel (x2 test, P < .05). Both use more quadrupedalism and climbing and less leaping. In P. melalophos the use of bimanual progression during feeding is negligible, and hopping is more common. Individuals of P. melalophos tend to hop between different parts of a large feeding tree, and climb or move quadrupedally within smaller regions of the tree. For all locomotor patterns, bout distances are significantly smaller in locomotion associated with feeding than in travel (t test, P < .05). The scale of distances involved in feeding sessions is smaller than that in travel; the former takes place within a tree, the latter within a home range. One can calculate from the percentage of bouts and the average bout distance, the contribution of each pattern of locomotion in an average kilometer of progression (Table 1, Figure 9). Since cost of lo- comotion for a


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