. The Compleat cladist : a primer of phylogenetic procedures. Cladistic analysis; Zoology -- Classification; Phylogeny. Fig. —Tree for ACCTRAN optimization (Exercise ). EXERCISE —Use Fig. for optimization and label the root node "0. 10 0 10 0 1. Fig. —^Tree for ACCTRAN optimization (Exercise ). Discussion ACCTRAN is a special case of a more general way to optimize characters on a tree (Swofford and Maddison, 1987). It will always insure that the length of the tree is minimized given a particular character matrix, but it will usually only show one interpretation of


. The Compleat cladist : a primer of phylogenetic procedures. Cladistic analysis; Zoology -- Classification; Phylogeny. Fig. —Tree for ACCTRAN optimization (Exercise ). EXERCISE —Use Fig. for optimization and label the root node "0. 10 0 10 0 1. Fig. —^Tree for ACCTRAN optimization (Exercise ). Discussion ACCTRAN is a special case of a more general way to optimize characters on a tree (Swofford and Maddison, 1987). It will always insure that the length of the tree is minimized given a particular character matrix, but it will usually only show one interpretation of how the characters evolved (a formal proof of Farris's algorithm is provided by Swofford and Maddison [1987]). That is, it favors reversals over repeated origins when the choice is equally parsimonious. We can see this by examining the very simple tree in Fig. When you optimize the characters on the tree using ACCTRAN, the result shows a reversal from 1 to 0 (Fig. b) (if this is not obvious, apply the ACCTRAN rules). The tree has a length of two steps. However, there is another equally parsimonious tree that interprets the evolution of a character coded "1" as a homoplasy (Fig. ). The problem is ACCTRAN will never allow you to find this Please note that these images are extracted from scanned page images that may have been digitally enhanced for readability - coloration and appearance of these illustrations may not perfectly resemble the original Wiley, E. O. Lawrence, Kan. : Museum of Natural History, University of Kansas


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