. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. THYIACINUS CYNOCEPHALUS Fig. 5: Tasmanian wolf, Thylacinus cynocephalus (Thylacinidae). Right ramus with incisors and canine (Tasmania). Redrawn from Hershkovitz (1982). Fig. 4 (left): Extinct borhyaena, Sipalocyon gracilis. Left ramus with incisors and canine: Top, labial view; bottom, Ungual view (Santacrucian, Argentina). Redrawn from Hershkovitz (1982). Amphyproviverra {Sipalocyon), and the caenolestid Halmarhiphus. As described by Sinclair (1906, p. 348, pis 40


. Bonner zoologische Beiträge : Herausgeber: Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn. Biology; Zoology. THYIACINUS CYNOCEPHALUS Fig. 5: Tasmanian wolf, Thylacinus cynocephalus (Thylacinidae). Right ramus with incisors and canine (Tasmania). Redrawn from Hershkovitz (1982). Fig. 4 (left): Extinct borhyaena, Sipalocyon gracilis. Left ramus with incisors and canine: Top, labial view; bottom, Ungual view (Santacrucian, Argentina). Redrawn from Hershkovitz (1982). Amphyproviverra {Sipalocyon), and the caenolestid Halmarhiphus. As described by Sinclair (1906, p. 348, pis 40, 45, fig. 3) the lower incisors of the Borhyaena "are closely crowded and the root of the second [is] is displaced posteriorly with reference to the median and lateral teeth, as in Thylacinus (fig. 4) and the Santa Cruz genera [of marsupials] in general!' Excluded were the Microbiotheriidae. Their lower in- cisors, Sinclair (1906, p. 409) noted, are "spatulate in shape, resembling the incisors of Dasyurus rather than Didelphis. UnHke these genera the root of the second tooth [is] in the series is not displaced posteriorly with reference to the roots of the first and third (fig. 5)!' See also Hershkovitz (1992). Morphological conditions for persistence of a staggered is have disappeared in the extinct South American Groberiidae and Argyrolagidae. The status of is in other ex- tinct American marsupials (Carolameghiniidae, Polydolopidae, and Patagoniidae) is unknown. Staggered is in Australian marsupials. AustraHan polyprotodont marsupials have lost an additional lower incisor, but the staggered is persists nonetheless in most. The incisor formula is 4/3 in the Dasyuridae (fig. 6), Thylacinidae (Order Dasyuromorphia, Marshall et al. 1990), all with staggered is. The incisor formula is 5/3 in the Peramelidae, Thylacomyidae and Notoryctidae (Order Peramelina, Mar- shall et al. 1990). In these, and the Myrmecobiidae, mandibular elongation coupled with attenuation tends to


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