. Asiatic herpetological research. Reptiles -- Asia Periodicals; Amphibians -- Asia Periodicals. Vol. 5, pp. 74-84 Asiatic Herpetological Research December 1993 Sympatric Amphibians of the Yew-box Grove, Caucasian State Biosphere Reserve, Sochi, Russia BORIS. S. TUNIYEV AND SVETLANA. Yll. BEREGOVAYA Caucasian State Biosphere Resen>e, Sochi, Russia Yew-box Grove of the Caucasian State Biosphere Reserve is home to seven species of amphibian. These species occur in a wide range of aquatic environments. The species composition, physical characteristics and history of each aquatic
. Asiatic herpetological research. Reptiles -- Asia Periodicals; Amphibians -- Asia Periodicals. Vol. 5, pp. 74-84 Asiatic Herpetological Research December 1993 Sympatric Amphibians of the Yew-box Grove, Caucasian State Biosphere Reserve, Sochi, Russia BORIS. S. TUNIYEV AND SVETLANA. Yll. BEREGOVAYA Caucasian State Biosphere Resen>e, Sochi, Russia Yew-box Grove of the Caucasian State Biosphere Reserve is home to seven species of amphibian. These species occur in a wide range of aquatic environments. The species composition, physical characteristics and history of each aquatic site was evaluated. The reproductive biology and food habits of each species was studied. These amphibians divide their niches on daily activity, seasonal activity, breeding site, microhabitat and food habits. The highest amphibian diversity and species overlap occurs in the most stable aquatic environments. Key Words: Amphibia, Russia, Caucasus, ecology. Introduction It is important when studying the influence of environmental factors on life history characteristics to distinguish those factors that are significant and those that are part of the "neutral background" (Monchadsky, 1958). The study of environmental influences is accomplished on sympatric species, usually closely related species (Orr and Maple, 1978; Ananjeva, 1981) at the population level (Pianka, 1973; Schoener, 1974; Schoener, 1977; Lyapkov and Severtsev, 1981). This is the study of the ecological niche (Pianka, et al., 1979. In the former Soviet Union one of the areas of highest amphibian diversity is found in the western Caucuses. The Yew- box Grove of the Caucasian State Biosphere Reserve is inhabited by eight amphibian species: Triturus vulgaris lantzi, T. cristatus karelini, T. vittatus ophryticus, P elodyte s caucasicus, Bufo verrucosissimus, Hyla arborea schelkownikowi, Rana ridibunda, and R. macrocnemis. Methods Field studies were conducted from 1980- 1982 in the Yew-box Grove (approximate are
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