. Bulletin of the British Museum (Natural History) Entomology. 88 IAN J. KITCHING Autographa Desertoplusia Diachrysia Macdunnoughia Antoculeora Plusia Plusidia Euchalcia Pseudochalcia Anadevidia Lamprotes Panchrysia Polychrysia Cornutiplusia Syngrapha Caloplusia Trichoplusia Argyrogramma Chrysodeixis Zonoplusia Ctenoplusia Acanthoplusia Erythroplasia Abrostola Fig. 5 Cladogram of the plusiine genera according to Kljutshko. (Redrawn from Kljutshko, 1985a, b.) blade-like setae on the valve), others were distinguished and delimited using symplesiomor- phies. For example, Autographa and Desertoplu
. Bulletin of the British Museum (Natural History) Entomology. 88 IAN J. KITCHING Autographa Desertoplusia Diachrysia Macdunnoughia Antoculeora Plusia Plusidia Euchalcia Pseudochalcia Anadevidia Lamprotes Panchrysia Polychrysia Cornutiplusia Syngrapha Caloplusia Trichoplusia Argyrogramma Chrysodeixis Zonoplusia Ctenoplusia Acanthoplusia Erythroplasia Abrostola Fig. 5 Cladogram of the plusiine genera according to Kljutshko. (Redrawn from Kljutshko, 1985a, b.) blade-like setae on the valve), others were distinguished and delimited using symplesiomor- phies. For example, Autographa and Desertoplusia were united, and separated from Diachrysia on the basis of a long harpe, the plesiomorphic state for his character 22. Similarly, the members of the 'clade' Autographa-Antoculeora were distinguished from Plusia by the presence of biordinal crotchets on the larval prolegs (Kljutshko's character 6). However, all other known plusiine larvae have biordinal crotchets and it is more parsimonious (and in keeping with previous authors' conclusions) to treat the uniordinal condition in Plusia as autapomorphic for this genus. This would appear to be a failure to employ the outgroup criterion correctly. If character 6 is reinterpreted and all the groups characterized by symplesiomorphies collapsed to form polytomies, then a much less well-resolved cladogram results (Fig. 6). Three tribes were recognized by Kljutshko: the Abrostolini, the Argyrogrammatini (including Erythroplusia) and the Plusiini. The last consisted of three subtribes: the Plusiina (including Anadevidia), the Caloplusiina and the Polychrysiina. Of the tribes, the Abrostolini and the Argyrogrammatini were sister-groups, in contrast to every other phylogeny that had been previously proposed for the Plusiinae. This relationship was based upon the possession of a. Please note that these images are extracted from scanned page images that may have been digitally enhanced for readability - coloration and appearance of these
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