. Bulletin of the Museum of Comparative Zoology at Harvard College. Zoology. LioPHOLiDOPHis (Colubridae) FROM MADAGASCAR • Cadle 439. Figure 34. Hemipenis of Liopholidophis epistibes, new spe- cies. Fully everted organ of MCZ 180318 (from Talatakely in the RNP), shown in sulcate (top) and asulcate (bottom) views. The distal "umbelliform" tips to the lobes in the sulcate view appear to be normal features, rather than a result of incomplete eversion (see text: "Summary and Comparisons of Hemipenes of Liopholidophis"). Scale bar = 1 mm. side the midsections bear 10-12 enlarged
. Bulletin of the Museum of Comparative Zoology at Harvard College. Zoology. LioPHOLiDOPHis (Colubridae) FROM MADAGASCAR • Cadle 439. Figure 34. Hemipenis of Liopholidophis epistibes, new spe- cies. Fully everted organ of MCZ 180318 (from Talatakely in the RNP), shown in sulcate (top) and asulcate (bottom) views. The distal "umbelliform" tips to the lobes in the sulcate view appear to be normal features, rather than a result of incomplete eversion (see text: "Summary and Comparisons of Hemipenes of Liopholidophis"). Scale bar = 1 mm. side the midsections bear 10-12 enlarged hooked spines (larger proximally, smaller distally) that grade into the small spines on the lobes. Large areas of nude tissue occupy the midsections between the clus- ters of spines. A few small spines are present in the fork of the sulcus. Small spines and spi- nules ornament the lobes except the nude central depression at their distal tips; a small wedge of nude tissue is in the crotch of the organ and adjacent basal facing por- tions of the lobes, forming a continuous stretch of nude tissue in the crotch between the asulcate and sulcate sides (, the spi- nous areas of the lobes are not continuous with one another across the crotch). As the lobes diverge from one another, their distal ends turn slightly toward the Figure 35. Hemipenis of Liopholidophis lateralis (Dumeril, Bi- bron, and Dumeril). Fully everted organ of MCZ 180380 (from near Midongy du Sud), shown in sulcate (top) and asulcate (bottom) views. The distal "umbelliform" tips to the lobes ap- pear to be normal features, rather than a result of incomplete eversion (see text: "Summary and Comparisons of Hemipenes of Liopholidophis"). Scale bar = 1 mm. asulcate surface, so that more of the tips of the lobes is visible from the asulcate than from the sulcate side. Lobes, except for the distal depression, are entirely ornamented with small spines, which are larger prox- imally. The distal tips
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