. A textbook of botany for colleges and universities ... Botany. BRYOPHYTES "3 initial. The mature archego- nium is stalked, with a massive venter, a long and often twisted neck, and numerous neck canal cells (fig. 249). Sporophyte.—The sporophyte (fig. 250) develops at first by a series of transverse walls until a short filament is formed, which later becomes massive by means of vertical walls. The upper tiers develop the capsule, which is organized as in Anthoceros, with the endothecium developed as the axial columella, with the sporogenous tissue cut ofif as an inner layer (becoming fo
. A textbook of botany for colleges and universities ... Botany. BRYOPHYTES "3 initial. The mature archego- nium is stalked, with a massive venter, a long and often twisted neck, and numerous neck canal cells (fig. 249). Sporophyte.—The sporophyte (fig. 250) develops at first by a series of transverse walls until a short filament is formed, which later becomes massive by means of vertical walls. The upper tiers develop the capsule, which is organized as in Anthoceros, with the endothecium developed as the axial columella, with the sporogenous tissue cut ofif as an inner layer (becoming four layers) of the amphithecium. Fig. 248. — Sphagnum: stem bearing a and capping the columella like terminal cluster of sporophytes, showing the a dome, and with a wall of five P°='''°" °^ archegonia. to seven layers. These Anthoceros-\ features are further emphasized by the large bulbous foot and the rudimentary seta, which is only a necklike connection between capsule and foot, and is often called the neck. Very important differences, however, are as follows: (i) there is no such development of chlorophyll tissue in the wall of the capsule as characterizes 4 n//ioce/'05; (2) the neck is not a growing region, and hence the capsule does not elongate ; and (3) the capsule dehisces by a definite lid {oper- culum), which is distinctly a moss character. As in all bryophytes, during the develop- ment of the sporophyte the venter of the archegonium develops the encasing calyptra, which in Sphagnum is ruptured by the. Please note that these images are extracted from scanned page images that may have been digitally enhanced for readability - coloration and appearance of these illustrations may not perfectly resemble the original Coulter, John Merle, 1851-1928; Barnes, Charles Reid, 1858-1910, joint author; Cowles, Henry Chandler, 1869- joint author. New York, Cincinnati [etc] American book company
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