. Asiatic herpetological research. Reptiles -- Asia Periodicals; Amphibians -- Asia Periodicals. Vol. 4, p. 90 Asiatic Herpetological Research February 1992. FIG. 9. Upper Pliocene changes of Phrynocephalus. A- middle Pliocene distribution of the species of the Phrynocephalus guttatus complex. The disjunction of the continuous distribution of this complex occured in the Upper Pliocene by: B- Junggar Alatau (Altai mountains); C- Saur and Tarbagatai mountain ridges. 1- Phrynocephalus melanurus; 2- Phrynocephalus versicolor ssp.; 3- Phrynocephalus versicolor paraskivi; 4- the place of speciation
. Asiatic herpetological research. Reptiles -- Asia Periodicals; Amphibians -- Asia Periodicals. Vol. 4, p. 90 Asiatic Herpetological Research February 1992. FIG. 9. Upper Pliocene changes of Phrynocephalus. A- middle Pliocene distribution of the species of the Phrynocephalus guttatus complex. The disjunction of the continuous distribution of this complex occured in the Upper Pliocene by: B- Junggar Alatau (Altai mountains); C- Saur and Tarbagatai mountain ridges. 1- Phrynocephalus melanurus; 2- Phrynocephalus versicolor ssp.; 3- Phrynocephalus versicolor paraskivi; 4- the place of speciation of the southwestern branch of the Phrynocephalus guttatus complex and Phrynocephalus moltschanovi. status of this species was discussed recendy by Golubev (1989b). It should be noted that the flow of Middle Asian rivers to the Caspian Sea changed to the Aral Sea, as a result of the downwarp of the region (Voskresensky, 1968). This was of crucial importance for the reconstruction of the ranges of most sclerobiont Phrynocephalus species. Probably before the Amu-Darya River (Uzboi), changed its course P. raddei was distributed on the clay and loess ares from the Caspian Sea to Kukhistan. Its range decreased considerably from the north and from the south owing to orogenesis (Fig. 9). In the south this species remained on the incline plain of the Kopeth Dagh and in the loess regions in the estuaries of the Murgab and Tedjen rivers. After the Amu Darya changed its flow to the Aral Sea, which coincided in time with the maximal development of the sand deserts, the area inhabited by P. raddei was divided into a number of isolated populations. These include the piedmont plains in Kukhistan and the Kopeth Dagh,. Please note that these images are extracted from scanned page images that may have been digitally enhanced for readability - coloration and appearance of these illustrations may not perfectly resemble the original Asiatic Herpetological Research Society; Chung-kuo liang chi
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