. Vegetation of Oregon and Washington. Botany Oregon Ecology; Botany Washington (State) Ecology. Figure 128. — Artemisia communities dominate the shrub-steppe of southeastern Oregon; in this High Lava Plains province landscape, Artemisia tridentata commu- nities occur on the hills, Artemisia arbuscula communities on the shallow soils of basalt flows, and Artemisia carta communities in seasonally ponded valley bottoms {photo courtesy Range Management, Oregon State Univer- sity). Columbia Basin province, are nearly absent. Hence, although the physiognomy and domi- nants of many communities are o
. Vegetation of Oregon and Washington. Botany Oregon Ecology; Botany Washington (State) Ecology. Figure 128. — Artemisia communities dominate the shrub-steppe of southeastern Oregon; in this High Lava Plains province landscape, Artemisia tridentata commu- nities occur on the hills, Artemisia arbuscula communities on the shallow soils of basalt flows, and Artemisia carta communities in seasonally ponded valley bottoms {photo courtesy Range Management, Oregon State Univer- sity). Columbia Basin province, are nearly absent. Hence, although the physiognomy and domi- nants of many communities are often similar, the environmental and community mosaics are different. The steppe and shrub-steppe communities of southeastern Oregon have not been com- prehensively treated. Nevertheless, from the generalized accounts of Poulton (1962), Anderson (1956), and Shantz and Zon (1924) and detailed but localized work by Eckert (1957), Dean (1960), Tueller (1962), Culver (1964), Dealy (1969), and McKell (1956), many of the important community types in the vegetational mosaic emerge. "ARTEMISIA" COMMUNITIES Artemisia communities dominate nearly every vegetational mosaic in southeastern Oregon's shrub-steppe (fig. 128). There are four major Artemisia species, each of which characterizes particular habitats: Artemisia tridentata on the deeper soils, Artemisia arbuscula on shallow, stony soils, Artemisia rigida on very shallow soils (Lithosols), and Artemisia cana on moister habitats. Associations of Artemisia tridentata, A. arbus- cula, and A. rigida have been described in de- tail (Eckert 1957; Culver 1964; HaU 1967); lists of characteristic species for six of these associations are provided in table 25. Artemisia tridentata/ Agropyron spicatum Association The Artemisia tridentata /Agropyron spica- tum association is very similar to the Colum- bia Basin province association of the same name (fig. 129); it is the most widespread as- Figure 129. — The Artemisia tridentata/Agro
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